Nemka Lelej, 1985

Lelej, Arkady S. & Williams, Kevin A., 2023, Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae), Zootaxa 5231 (2), pp. 101-140 : 118

13. Nemka Lelej, 1985

( Figs 41–44 , 115 )

Nemka Lelej, 1985: 240 , ♁, ♀ (as subgenus of Smicromyrme Thomson); 1995a: 2; Lelej & Brothers 2008: 39; Brothers & Lelej 2017: 95, ♁, ♀; Williams et al. 2019: 22, ♀; Pagliano et al. 2020: 169; 176; Okayasu 2020: 24, ♁, ♀.

Horaia Tsuneki, 1993: 43 (as subgenus of Smicromyrme Thomson ). Type species Smicromyrme (Horaia) chihpenchius Tsuneki, 1993 , ♁, by original designation. Junior homonym of Horaia Tonnoir, 1930 ( Diptera ). Junior subjective synonym of Nemka Lelej, 1985 according to Lelej, 1995a: 2.

Type species: Mutilla viduata Pallas, 1773 , ♁, by original designation.

Diagnosis. MALE. Head not elevated posterad, frons and vertex usually punctate, not longitudinally striate. Mandibles robust apically tridentate, deeply excised beneath with large subbasal tooth. Hypostomal bridge without tooth. Ocelli not enlarged. Scape obscurely bicarinate beneath. Mesopleuron with one or two precoxal denticles or tubercles beneath. Tegula rounded posterad, not or slightly protruding behind mesoscuto-scutellar suture. Wing dark fuscous, stigmatic cell slightly less than distance between base of stigmatic cell and origin of RS on vein Sc. Metasoma usually with some segments red or yellowish-red. T1 shorter than half of T2. T2 with longer lateral felt lines, S2 with short lateral felt lines. Volsellar cuspis long and narrow with long setae, volsella with basal external lobe and long setae; setae on inner and ventral margin of external lobe extremely long and curved. FEMALE. Unlike the other Oriental and Palaearctic Smicromyrmini , Nemka females have the pygidium well-defined and short oval in shape. Other useful characters include the presence of a distinct scutellar scale, a single medial setal spot on the T2 disc, and the posterior setal band of T2 variably expanded. Minimum distance between eyes more than 1.4 × eye height; genal carina and hypostomal tooth absent; clypeus with subapical transverse ridge; prementum without dome-like tubercle. Mandible slender. Pronotum without projecting humeral angle; scutellar scale distinct. Protarsus with external comb of long spines. T1 without pale setal spots; T2 with basomedial spot and apical spot or narrow band of pale setae, the latter slightly widened medially; T3 with wide uniform band of pale setae; pygidial plate short oval, distinctly carinate laterally; surface of pygidial plate longitudinally rugoso-striate throughout or unsculptured apically.

Diversity and distribution. Eighteen species are recognized: Nemka viduata ( Pallas, 1773) with five subspecies and N. aurantiaca ( Skorikov, 1935) are distributed in Palaearctic, and 16 species occur in Oriental Region, three of these penetrate to the Palaearctic parts of China and Iran. Seven species are known from both sexes, three are known from males only, and eight are known from females only.

Remarks. More than half of the known species are included in Lelej’s (1995a) key.

Figures

FIGURES 35–44. 35–40. Mickelomyrme hageni (Zavattari, 1913): 35–37. M. hageni, ♀ (=Mutilla alberici Zavattari, 1913, ♀, lectotype); 38–40. M. hageni, ♁ (Mutilla hageni Zavattari, 1913, ♁, lectotype). 41–44. Nemka viduata (Pallas, 1773), ♀ (41, 42), ♁ (43, 44). 35, 38, 41. Dorsal habitus; 36, 39, 42, 44. Lateral habitus; 37, 40. Labels; 43. Head, face view. (Photographs 35–37 from Integrated insect types database of Taiwanese species http://twinsecttype.nmns.edu.tw/specimen/?id=NMNS-SDEI- 01376; 38–40 http://twinsecttype.nmns.edu.tw/specimen/?id=NMNS-SDEI-01380) and by K. Williams (41–44)).

FIGURES 110–115. Smicromyrmini, ♀, pygidial plate (T6). 110.Andreimyrme rong Williams in Okayasu et al., 2021, holotype; 111. Dentilla curtiventris (André, 1901); 112. Ephucilla thalia (Mickel, 1933); 113. Ephutomma fletcheri (Turner, 1911); 114. Karunaratnea palatupanae Lelej, 2005, paratype; 115. Nemka viduata (Pallas, 1773). (Photographs by K. Williams andA. Lelej (114)).