Sinotilla Lelej, 1995

Lelej, Arkady S. & Williams, Kevin A., 2023, Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae), Zootaxa 5231 (2), pp. 101-140 : 125-126

22. Sinotilla Lelej, 1995

( Figs 74–76 )

Sinotilla Lelej, 1995b: 3 , 4 (key), 18 (description), ♁, ♀; Lelej & Brothers 2008: 55; Brothers & Lelej 2017: 96, ♁, ♀; Okayasu 2017: 152 , ♀; 2018: 486, ♀, ♁; Okayasu et al. 2018: 313 , ♀; Williams et al. 2019: 25 , ♀; Pagliano et al. 2020: 186; Okayasu 2020: 87 , ♁, ♀.

Type species Smicromyrme boheana Chen, 1957 , ♁, by original designation.

Diagnosis. MALE. Head robust, elevated posterad, frons and vertex longitudinally striate. Mandible bidentate at apex, deeply excised beneath, with large subbasal tooth. Clypeus weakly convex or weakly concave, with protruding midpart of anterior margin. Ocelli small, POD (in type species) 4.3 × posterior ocellus diameter; each ocellus posterad with longitudinal carina, running to occiput. Scape distinctly bicarinate beneath, emargination between carinae with setae. F1 (in type species) half its maximal width and 0.3 × F2. Tegula protruding beyond scutoscutellar suture for 1/5–1/4 of its length. Parascutal carinae absent. Notauli not visible (in type species) or invisible in anterior one-third (in S. hong Lelej, 1995 ). Parapsidal furrow short. Scutellum simple. Mesopleuron beneath with longitudinal, glabrous, precoxal line. Metacoxa mesally carinate. Wings infuscated; stigmatic cell is less than 0.6 × distance between base of stigmatic cell and origin of RS on vein Sc. T1 with distinct dorsum, T2 with lateral felt lines, S2 with shorter ones (in type species few small punctures only). T7 densely punctate (in type species) or with indefinite median impunctate area (in S. hong ). S8 without tubercles or carinae, densely punctate. Volsella without basal external lobe; volsellar cuspis short and oval, apically and its inner margin with setae not longer than cuspis. FEMALE. Mandible slender. Clypeus with protruding transverse lamellate carina and median basal tubercle. Fore tarsus without external comb of spines. Genal carina developed, with tubercle on hypostomal carina. F1 0.8–1.2 × its maximal width, 1.4–1.6 × pedicel, 1.0–1.5 × F2, latter slightly longer or shorter than F3. Mesosoma convergent posterad (not convergent posterad in S. belokobylskiji Lelej, 1995 ). Scutellar scale weak, but distinct. T2 apically with band of pale setae, slightly widened medially. T3 with wide pale band of pale setae. T6 with glabrous shiny medial part, not carinated laterad even apically, considerably convergent basally with basolateral tufts of dense pale setae.

Diversity and distribution. Twenty species (two subdivided into subspecies) are recognized from the Oriental Region, S. pekiniana ( André, 1905) penetrates to the Palaearctic part of China ( Pagliano et al. 2020), and S. jambar Okayasu 2020 is described from Okinawa ( Japan) ( Okayasu 2020). One is known from both sexes, nine are based on males only, and 10 are based on females only.

Remarks. Females of the genus were reviewed by Okayasu (2017), but many additional species have been treated or described subsequently (e.g. Okayasu et al. 2018, Okayasu 2020).

Figures

FIGURES 74–81. 74. Sinotilla boheana (Chen, 1957), ♁, lateral habitus; 75–76. S. belokobylskiji Lelej, 1995, ♀: 75. Dorsal habitus, 76. Lateral habitus. 77–79. Skorikovia elongata (Radoszkowski, 1885): 77. ♁, lateral habitus; 78. ♀, dorsal habitus; 79. ♀, lateral habitus; 80–81. Tsunekimyrme fluctuata (Smith, 1865), ♁: 80. Lateral habitus, 81. Head, face view. (Photographs by K. Williams and A. Lelej (74, 80, 81)).