Anomala robiginosa Filippini, Galante, and Micó, 2015

Filippini, V., Galante, E. & Micó, E., 2015, Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica, The Coleopterists Bulletin 69 (3), pp. 463-476 : 473

publication ID	https://doi.org/ 10.1649/0010-065x-69.3.463
publication LSID	lsid:zoobank.org:pub:9A0330AD-EEF7-4A80-8D80-1B6116BD856F
persistent identifier	https://treatment.plazi.org/id/03CFA348-6B6C-FFC9-FF7E-FE6F5981FAF4
treatment provided by	Diego
scientific name	Anomala robiginosa Filippini, Galante, and Micó
status	sp. nov.

Treatment

Anomala robiginosa Filippini, Galante, and Micó new species

( Figs. 6 View Figs , 14 View Figs , 22 View Figs , 30 View Figs , 34 View Figs )

Type material. Holotype: ♂ " Zarcero , Alfaro Ruiz, 1700m Alajuela Pr. COSTA RICA. Apr - May 1989. A. Del Valle & A. Solis, L- N 240500_493500 / INBIOCRI002517719" deposited at MNCR . Paratypes (3): 1♀ " Santa Rosa National Park Guana. Prov. Costa Rica 18–20 May 1979 D. H. Janzen / INBIOCRI000111925" ; 1♂ " ALAJUELA, COSTA RICA Zarcero , 1700 m 23-V-87 COL: A. SOLIS B. / CEUA00106202 " ; 1♂ " Alajuela. Costa Rica. Zarcero , 1700 m 23 may 1987 Angel Solís / INBIOCRI002517717". Two paratypes are deposited at MNCR , and one paratype is deposited at Universidad de Alicante .

Holotype. Male ( Fig. 6 View Figs ). Body oval. Length 11.23 mm, width 5.75 mm. Head, pronotum, and scutellum bronze-brown. Elytra ochre with suture and punctures reddish brown. Pygidium, legs, and venter orange-brown. Head: Clypeus trapezoidal, surface densely punctate-reticulate, convex. Anterior margin straight, thin in frontal view, width/ length ratio 2.09. Frons densely punctate, with a parabolic concavity. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.48. Ratio of antennal funiculus/club 0.7. Pronotum: Trapezoidal. Ratio width/length 1.84. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles obtuse. Basal margin sinuate, obliterated at middle. Surface with deep, dense punctures. Scutellum: Subtriangular, with straight lateral margins and acute apex. Ratio width/length 1.27. Surface densely punctate. Elytra: With costae defined by rows of sparse, coalescing punctures. Subsutural interstice with 1–2 rows of sparse punctures, 2 nd –4 th interstices with 1 irregular, secondary stria. Pygidium: Finely granulated, with long setae on posterior margin. Ratio width/length 1.65. Venter: Space between mesocoxae narrow, slightly convex. Mesometasternal suture welldefined at base of mesocoxae. Abdominal sternites with 1 row of setae (2 rows on second sternite) and 3–4 rows of punctures per sternite. Last sternite punctate, with subapical bead thick and sinuate, apical membrane wide. Legs: Protibia ( Fig. 14 View Figs ) bidentate, apical tooth long and straight, basal tooth below internal apex of protibia triangular, sharp, obtuse. Metatibia slightly narrower sub- apically. Ratio length/width 2.98. First external carina with row of sparse setae. Surface punctate above second external carina and rugose below. Internal protarsal claw bifurcate, with upper branch slightly shorter and 1/3 width of lower one. Inferior margin with a sharp angle at half length. Male genitalia: Parameres slender ( Fig. 22 View Figs ), inferior margin sinuate. Apical portion of ventral plate membranous. Endophallus ( Fig. 30 View Figs ) developing dorsally with respect to median lobes; sclerotized, flat plate present posteriorly at base, above which a rounded, hairy inflation is present; ejaculatory duct opening located frontally.

Female. Similar to male except protibia ( Fig. 14 View Figs ) with longer and wider first tooth; internal protarsal claw narrower with curve inferior margin; last sternite with curved apex.

Variation. Three paratypes. Head, pronotum, and venter vary from orange to dark brown; presence of small dark maculae on basal third of elytra. Body length 10.68–11.49 mm, body width 5.75– 6.44 mm. Clypeal width/length ratio 2.01–2.33. Ratio of interocular width/width of eye 2.24–2.75. Ratio of antennal funiculus/club 0.67–0.74. Pronotal width/length ratio 1.72–1.84. Scutellar width/length ratio 1.24–1.37. Pygidial width/length ratio 1.65–1.82. Metatibial length/width ratio 2.77–3.03.

Etymology. From the Latin adjective robiginosus, -a, -um, rusty, for the rust-colored elytral maculae.

Distribution. Anomala robiginosa inhabits the dry forest in the northwestern Guanacaste lowlands and Tilarán and northwestern extreme of Central mountain ranges of Costa Rica, from 500 to 1,700 m elevation ( Fig. 34 View Figs ).

Diagnosis. Pronotum bronze-brown, elytra with pigmented punctures, parameres slender, apex squared in lateral view, endophallus developing dorsally with respect to the median lobes. This species is very similar to A. sticticoptera , but it can be distinguished by the shape of the aedeagus. In A. sticticoptera , the parameres are shorter, with a narrow apex, and the ventral angle is acute and protruding posteriorly ( Fig. 33 View Figs ).