Anemesia sogdiana, Zonstein, 2018
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publication ID |
https://doi.org/ 10.5852/ejt.2018.485 |
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publication LSID |
lsid:zoobank.org:pub:55A0F74D-FA80-4C6A-AD74-B49C9061A449 |
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DOI |
https://doi.org/10.5281/zenodo.3848281 |
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persistent identifier |
https://treatment.plazi.org/id/15FFF793-3F27-4659-BE0D-15674535BD7A |
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taxon LSID |
lsid:zoobank.org:act:15FFF793-3F27-4659-BE0D-15674535BD7A |
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treatment provided by |
Valdenar |
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scientific name |
Anemesia sogdiana |
| status |
sp. nov. |
Anemesia sogdiana View in CoL sp. nov.
urn:lsid:zoobank.org:act:15FFF793-3F27-4659-BE0D-15674535BD7A Figs 7 View Figs 1–9 , 20 View Figs 18–26 , 30 View Figs 27–33 , 54 View Figs 48–62 , 68–69 View Figs 63–77 , 82 View Figs 78–89 , 96 View Figs 90–98 , 109 View Figs 108–116 , 124 View Figs 117–124 , 133–137 View Figs 133–142 , 154–155 View Figs 143–157 , 165–168 View Figs 158–172 , 179 View Figs 173–187 , 203–205 View Figs 197–205 , 235–240 View Figs 224–244 , 277–278 View Figs 266–277 View Figs 278–290 , 295–297, 314–316, 332, 369
Brachythele birulai – Charitonov 1969: 66 (Ƌ). Misidentified, not Brachythele birulai Spassky, 1937 .
Diagnosis
In habitus, Anemesia sogdiana sp. nov. is largely similar to A. pococki sp. nov. and especially to A. andreevae sp. nov., but differs from them by the long, thin and gradually curved embolus in males ( Figs 203–205 View Figs 197–205 , cf. Figs 193–194 View Figs 188–196 , 197–198 View Figs 197–205 ) as well as by the entire and narrowed subapically spermathecae in females ( Figs 235–240 View Figs 224–244 , cf. Figs 228–229, 233–234 View Figs 224–244 ).
Etymology
Sogdiana (‘ ΣΟΓΔΙΑνή ’, an ancient Greek derivate from the Old Persian Suguda) is the name of an historical area in Central Asia that existed in the late antiquity and in the early Middle Ages (IVth century BC – IXth century AD) and is applied to the territory located between the Amu-Darya and Syr-Darya Rivers which corresponds to the known range of this species. This specific epithet is a noun in apposition to the genus name, the gender is feminine.
Material examined
Holotype
UZBEKISTAN: Ƌ, northwestern tip of Zeravshan Mts , Amankutan Pass, 1670 m, 39°17′27′′ N, 66°54′01′′ E, 5 May 1995, S. Zonstein leg. ( SMNH).
GoogleMapsParatypes (4 ƋƋ, 17 ♀♀)
UZBEKISTAN: 1 Ƌ, same collection data as for the holotype ( SMNH); 1 Ƌ, same locality, 1650– 1750 m, 39°18′ N, 66°54′ E, 11 Apr. 1966, V.F. Bakhvalov leg. ( SMNH); 1 ♀, same collection data as for preceding but 7 Apr. 1989, S. Zonstein leg. ( SMNH); 1 Ƌ, 9 ♀♀, same collection data as for preceding but 27 Apr. 1993 ( SMNH); 2 ♀♀, Zeravshan Mts, foothills 10 km N of Kitab, 1000 m, 39°12′ N, 66°54′ E, 9 Apr. 1989, S. Zonstein leg. ( SMNH); 1 Ƌ, Zeravshan Mts, Jindy-Daria Canyon, Hojakurgan, 1400–1800 m, 39°11′ N, 67°17′ E, 14 Apr. 1990, S. Zonstein leg. ( SMNH); 2 ♀♀, same collection data as for preceding but 9 Apr. 1991 ( SMNH); 2 ♀♀, same collection data as for preceding but 26 Apr. 1992 ( SMNH).
Additional material (3 ƋƋ, 45 ♀♀, 2 ♀♀ subad.)
UZBEKISTAN: 2 ƋƋ, northwestern slope of Hissar Mts, surroundings of Ishkent, 1100–1300 m (38°51′ N, 66°58′ E), 25–28 Mar. 1942, K. Arnoldi and D. Fedotov leg. ( PSU); 1 Ƌ, Turkestan Mts, foothills near Zaamin (700 m, 39°57′ N, 68°23′ E), 8Apr. 1980, F. Khassanov leg. ( SMNH); 2 ♀♀ subad., northwestern ending of Nuratau Mts, Zafarabad (Kokcha), 400 m, 40°32′ N, 65°02′ E, 7 Apr. 1990, S. Zonstein leg. ( SMNH); 32 ♀♀, Baisuntau Mts, 2 km E Shurob, 1050–1200 m, 38°12′ N, 67°00′ E, 15 Apr. 1987, S. Zonstein leg. ( SMNH, ZMMU); 3 ♀♀, Hissar Mts, Majanak 10 km N Kokbulak, 2500– 2650 m, 38°41′ N, 66°56′ E, 6 Jun. 1997, S. Zonstein leg. ( SMNH); 3 ♀♀, Kugitang Mts, Baglydara canyon, 1300–1800 m, Jul. 1983, A.B. Nenilin leg. ( SMNH); 4 ♀♀, same locality, 37°53′ N, 66°40′ E, 1700 m, 15 May 1985, S. Zonstein leg. ( SMNH); 1 ♀, same locality, 8 Apr. 1989, S. Zonstein leg. ( SMNH); 2 ♀♀, Kugitang Mts, canyon 3 km SW of Vandob, 1600–1700 m, 37°42′ N, 66°34′ E, 30 May 1995, S. Zonstein leg. ( SMNH).
TAJIKISTAN: 3 ♀♀, southern slope of Hissar Mts, foothills 8 km N Dushanbe, Varzob canyon, 1100 m, 38°40′ N, 68°47′ E, 25 May 2002, S. Zonstein leg. ( SMNH).
Description
Male (holotype)
HABITUS. See Fig. 7. View Figs 1–9
MEASUREMENTS. TBL 13.40, CL 5.15, CW 4.53, LL 0.60, LW 0.99, SL 2.86, SW 2.28.
COLOUR. Carapace light yellowish brown with anterior edge darker and thoracic part paler; chelicerae, majority of palps and legs light yellowish brown; sternum, labium, maxillae and leg tarsi paler; eye tubercle blackened; abdomen dorsally yellowish grey with brown pattern consisting of moderately wide median longitudinal spot and few paired transverse and slightly inclined short stripes, ventral part of abdomen pale yellowish grey, spinnerets pale brownish yellow.
PROSOMA. Clypeus and eye tubercle as in Fig. 54 View Figs 48–62 . Eye diameters and interdistances: AME 0.15(0.21), ALE 0.30, PLE 0.15, PME 0.06, AME–AME 0.15(0.09), ALE–AME 0.15(0.12), ALE–PLE 0.11, PLE– PME 0.05, PME–PME 0.48. Weak cheliceral rastellum composed of several long bristles lacking tips and grouped in one transverse row in front of fang base and in rastellar mound. Each cheliceral furrow with 8–9 promarginal teeth and 0–1 small retrobasal teeth. Sternum, labium and maxillae as shown in Fig. 96 View Figs 90–98 . Sternal sigilla small, posterior pair long-oval and distant from sternum edge. Maxillae with 7–9 cuspules each.
LEGS. Tibia and metatarsus I as in Fig. 124 View Figs 117–124 . Scopula: entire and distal on metatarsi I and II, narrowly divided on tarsi I and II, mixed and widely divided on tarsus III, vestigial on tarsus IV. Trichobothria: two rows of 8–9 each on tibiae, 11–13 on metatarsi, 12–14 on tarsi, 10 on cymbium. PTC I–II: outer and inner margins with 6–7 teeth each. PTC III with 6–7 and 5–6; PTC IV with 6 and 3, respectively.
SPINATION. Palp: femur d3, pd1; patella pd1; tibia p3, pv1; tarsus d7–8. Leg I: femur d5, pd3; patella pd1; tibia pd3, p1, r3, v6–7+m; metatarsus d1, p2, v1. Leg II: femur d5, pd3; patella p1; tibia p3, v8; metatarsus d2, p4, v7. Leg III: femur d4, p3, r2; patella p3, r1; tibia d1, p3, r3, v7; metatarsus d2, p4, r3, v8; tarsus p1–2. Leg IV: femur d5, r1; tibia d1, p3, r3, v7; metatarsus p4, r4, v8–9; tarsus p1. Patella IV and tarsi I–II unarmed.
PALP. Tibia, cymbium and palpal organ as shown in Figs 179 View Figs 173–187 , 204–205 View Figs 197–205 . Tibia relatively short, swollen, and aspinose ( Fig. 179 View Figs 173–187 ). Palpal organ with long, tapering and slightly curved embolus ( Figs 204–205 View Figs 197–205 ).
SPINNERETS. See Fig. 277 View Figs 266–277 . PMS: length 0.44, diameter 0.16. PLS: maximal diameter 0.54; length of basal, medial and apical segments 0.92, 0.70, 0.67; total length 2.29; apical segment short-digitiform.
Female (paratype from Amankutan)
HABITUS. See Fig. 20. View Figs 18–26
MEASUREMENTS. TBL 16.50, CL 6.93, SW 5.38, LL 0.87, LW 1.33, SL 3.30, SW 2.97.
COLOUR. Carapace and legs dorsally light yellowish foxy brown; cephalic part darkened, medium foxy brown; eye tubercle with blackish brown spots around AMEs and lateral eyes; chelicerae reddish brown; sternum, labium, maxillae and legs ventrally pale brownish yellow; abdomen light greyish brown with dark brown dorsal pattern consisting of wide median longitudinal spot and six pairs of lateral chevrons; spinnerets very pale greyish brown.
PROSOMA. Clypeus and eye tubercle as in Fig. 68 View Figs 63–77 . Eye diameters and interdistances: AME 0.18(0.24), ALE 0.38, PLE 0.20, PME 0.15, AME–AME 0.18(0.12), ALE–AME 0.20(0.17), ALE–PLE 0.15, PLE– PME 0.04, PME–PME 0.52. Cheliceral rastellum composed of about 30 spikes located mostly on low mound. Each cheliceral furrow with 8–9 promarginal teeth and one smaller retrobasal tooth. Sternum, labium and maxillae as shown in Fig. 109 View Figs 108–116 . Sternal sigilla minute and oval, posterior pair long-oval and distant from sternum edge. Maxillae with 14–16 cuspules each.
SPINATION. Palp: femur pd2; patella p1; tibia p5, v10–12; tarsus v2–3. Leg I: patella p3; tibia v7–8; metatarsus p1, v8–9. Leg II: patella p1; tibia p3, v7–8; metatarsus p2–3, v9–10. Leg III: patella p3; tibia d1; p1, r1, v7; metatarsus pd3, p4, dr3, v7; tarsus p2, v1. Leg IV: tibia v9–10; metatarsus p1, r1, v8; tarsus v2. All femora dorsally with 4–6 true spines and several bristles; patella IV and tarsi I and II aspinose.
LEGS. See Figs 133–137 View Figs 133–142 . Scopula entire and distal on metatarsi I and II, entire on palpal tarsus, narrowly divided on tarsi I and II, elsewhere absent. Trichobothria: two rows of 7–9 each on tibiae, 10–12 on metatarsi, 12–14 on tarsi, 10 on palpal tarsus. Palpal claw with 4 promarginal teeth. PTC I–II with 4–5 teeth on each margin. PTC III with 4–5 teeth on outer, and 2–3 teeth on inner margins; PTC IV with 3–4 and 1–2 teeth, respectively. Difference in dentition between PTC I and PTC IV as shown in Figs 154–155. View Figs 143–157
SPERMATHECAE. Moderately long, entire, and narrowed apically. See Fig. 236. View Figs 224–244
SPINNERETS. See Fig. 278 View Figs 278–290 . PMS: length 0.62, diameter 0.32. PLS: maximal diameter 0.62; length of basal, medial and apical segments 0.98, 0.70, 0.63; total length 2.33; apical segment triangular. Spigots as shown in Figs 295–297 View Figs 291–305 .
Variation
The length of the carapace varies from 4.87 to 5.21 in males and from 4.47 to 7.33 in females. Colouration varies through specimens very narrowly (see Fig. 30 View Figs 27–33 ). Throughout the specimens, both PLE and PME may be reduced or even lost ( Fig. 69 View Figs 63–77 ). In some females, the cheliceral furrow may be armed with 2–3 retromarginal teeth (see Fig. 82 View Figs 78–89 ). In conspecific males, the shape of the palpal organ is almost identical ( Figs 203–205 View Figs 197–205 ). Spermathecae show some variations and can feature with straight or bent stalks ( Figs 235–240 View Figs 224–244 ).
Habitat
The known records mostly correspond to the midland mountain belt although the species can also occur in both the piedmont semi-desert (Zafarabad) and the highlands (Majanak); the preferred biotopes are represented by steppes and open forest communities with different species of Juniperus L., Crataegus Tourn. ex L. and Acer L. (see Figs 314–316 View Figs 314–321 ).
Distribution
Central and southern Uzbekistan, and western Tajikistan ( Fig. 369).
Note
Charitonov (1969) listed five males of this species collected by K. Arnoldi and D. Fedotov in 1942 in the surroundings of Ishkent as Brachythele birulai Spassky. A detailed examination of the spider collection at the Perm University (1987, 1988) revealed the presence of only two appropriate male specimens sampled and labelled according to the mentioned record data.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Anemesia sogdiana
| Zonstein, Sergei 2018 |
Brachythele birulai
| Charitonov D. E. 1969: 66 |