Anemesia castanea, Zonstein, 2018
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Figs 8 View Figs 1–9 , 21 View Figs 18–26 , 55 View Figs 48–62 , 70 View Figs 63–77 , 97 View Figs 90–98 , 110 View Figs 108–116 , 125 View Figs 125–132 , 180 View Figs 173–187 , 206–208 View Figs 206–214 , 245 View Figs 245–265 , 279 View Figs 278–290 , 318–321, 371 View Fig
Anemesia castanea sp. nov. resembles A. karatauvi in habitus, as well as in possessing the similar looking comma-shaped palpal organ provided with a long, thin and gradually curved embolus in males, and relatively long and slender tapering spermathecae in females. However, in comparison with the latter species, males of A. castanea sp. nov. have a more slender male palpal tibia, a domed ventral surface of the palpal bulb and a longer embolus (vs stouter tibia, flattened surface, and shorter embolus; Figs 180 View Figs 173–187 , 206–208 View Figs 206–214 , cf. Figs 181 View Figs 173–187 , 209–210 View Figs 206–214 ). Females of A. castanea sp. nov. differ from those of A. karatauvi in having the closer spaced and more diverged spermathecae ( Fig. 245 View Figs 245–265 , cf. Figs 246–249 View Figs 245–265 ), as well as by a larger number of the maxillary cuspules (65–70 vs 40–55) and by a better developed cheliceral rastellum.
The specific name is a derivative (using the feminine gender) of the Latin adjective ‘ castaneus ’ meaning ‘chestnut-brown’; the name is derived from the colouration of the specimens.
TAJIKISTAN: Ƌ, foothills of Aruktau Mt. R., 0.5 km SE of Ganjina Village, 750 m, 37°58′ N, 68°32′ E, 14 Apr. 1991, S. Zonstein leg. ( SMNH).GoogleMaps
TAJIKISTAN: 4 ƋƋ same collection data as for the holotype but S. Zonstein and S. Ovchinnikov leg. ( SMNH).
TAJIKISTAN: 1 ♀, Ganjina, under stone, 9 Apr. 1967, E. Andreeva leg. ( MIZW), 1 ♀ (dried), Lower Vahsh Valley, Tigrovaya Balka (Tiger Gorge) Reserve, 12–18 Apr. 1967, E. Andreeva leg. ( MIZW). See note below.
HABITUS. See Fig. 8. View Figs 1–9
MEASUREMENTS. TBL 15.90, CL 6.15, CW 5.54, LL 0.56, LW 1.07, SL 2.96, SW 2.77.
COLOUR. Carapace almost uniformly light chestnut brown; chelicerae, leg I entirely and most part of palps and legs II–IV light-medium to dark brown; sternum, labium, maxillae and leg tarsi paler, brownish orange; eye tubercle black; abdomen dorsally yellowish grey with dark brown pattern consisting of dentate median stripe anteriorly and few paired transverse and slightly inclined chevrons posteriorly; ventral part of abdomen and spinnerets pale yellowish grey.
PROSOMA. Clypeus and eye tubercle as in Fig. 55 View Figs 48–62 . Eye diameters and interdistances: AME 0.15(0.23), ALE 0.25, PLE 0.22, PME 0.18, AME–AME 0.21(0.13), ALE–AME 0.12(0.08), ALE–PLE 0.12, PLE– PME 0.02, PME–PME 0.47. Cheliceral rastellum consists of about 20–25 spikes in front of fang base and on very low mound. Each cheliceral furrow with 7 promarginal teeth and 5–7 smaller retromarginal teeth. Sternum, labium and maxillae as shown in Fig. 97 View Figs 90–98 . Sternal sigilla small oval, posterior pair considerably distant from sternum edge. Maxillae with ca 35 cuspules each.
LEGS. Tibia and metatarsus I as in Fig. 125 View Figs 125–132 . Scopula: distal and entire on metatarsi I and II, narrowly divided on tarsi I and II, widely divided on tarsi III, absent on tarsi IV. Trichobothria: two rows with 9–11 each on tibiae, 20–24 on metatarsi, 16–20 on tarsi, 13–14 on cymbium. PTC I–II with 11–13, PTC III–IV with 9–10 teeth on each margin, respectively.
SPINATION. Palp: femur d4, pd4, rd1; patella p1; tibia d1, p3–4, pv3, rv7; tarsus d6–7. Leg I: femur d4, pd3–4, rd3; tibia p3–4, r2, v9+m; metatarsus pd3, v5–7. Leg II: femur d4, pd3–4, rd3–4; tibia p3, v7–8; metatarsus p4, v5–6; tarsus p2. Leg III: femur d4, pd3, rd3; patella p3, r2; tibia d2–3, p3, r3, v7; metatarsus dp4, r3–4, v7–8; tarsus p2–4. Leg IV: femur d4, p3, r2–3; patella r1; tibia d1, p3, r3, v8–9; metatarsus d6–7, p5, r4–5, v9–10; tarsus p2, r2. Patellae I, II, and tarsus I aspinose.
PALP. Tibia, cymbium and palpal organ as shown in Figs 180 View Figs 173–187 , 207–208 View Figs 206–214 . Palpal tibia very long, slender and cylindrical, with spines and numerous stout bristles ( Fig. 180 View Figs 173–187 ). Palpal organ with embolus very long, tapering and slightly curved ( Figs 207–208 View Figs 206–214 ).
SPINNERETS (see Fig. 279 View Figs 278–290 ). PMS: length 0.46, diameter 0.20. PLS: maximal diameter 0.57; length of basal, medial and apical segments 0.94, 0.68, 0.76; total length 2.38; apical segment shortly digitiform.
Female (from Ganjina)
HABITUS. See Fig. 21. View Figs 18–26
MEASUREMENTS. TBL 25.20, CL 8.39, CW 7.44, LL 1.01, LW 1.60, SL 4.26, SW 4.16.
COLOUR. As in male.
PROSOMA. Clypeus and eye tubercle as shown in Fig. 70 View Figs 63–77 . Eye diameters and interdistances: AME 0.15(0.25), ALE 0.32, PLE 0.24, PME 0.26, AME–AME 0.36(0.26), ALE–AME 0.28(0.23), ALE–PLE 0.27, PLE–PME 0.01, PME–PME 0.83. Cheliceral rastellum represented by approximately 40 spikes located in front of fang base and on low mound. Each cheliceral furrow with 9 promarginal teeth and 10–11 smaller retromarginal teeth. Sternum, labium and maxillae as shown in Fig. 110 View Figs 108–116 . Sternal sigilla small; posterior pair distant from sternum edge, narrowly oval. Maxillae with ca 70 cuspules each.
LEGS. Scopula dense, occupies 5/6 ventral surface of metatarsus I, distal on metatarsus II, entire on palpal tarsus and tarsus I, widely divided on tarsus II, elsewhere absent. Palpal claw with 5 promarginal teeth. Trichobothria: 2 rows of 10–13 each on tibiae, 14–17 on metatarsi, 13–15 on leg tarsi, 13 on palpal tarsus. PTC I–II with 6–7 teeth on each margin. PTC III with 5–6 teeth on outer, 4–5 teeth on inner margins; PTC IV with 4–5 and 0–4 teeth, respectively.
SPINATION. Palp: femur pd1; patella p2; tibia p2, v14–15; tarsus v2. Leg I: femur pd1; patella p1; tibia v5; metatarsus v5. Leg II: femur pd2; patella p2; tibia p2, v5; metatarsus p2, v5–6. Leg III: femur pd2, rd3; patella p2; tibia d1; p3, r2, v7; metatarsus pd2, p3, rd3, v8–9; tarsus p1, v2. Leg IV: femur rd2; tibia r1, v6; metatarsus pd1, p2, rd2, v7–11; tarsus v2. All femora with 1 basodorsal spine and dorsal row of few bristles. Patella IV, and tarsi I and II aspinose.
SPERMATHECAE. Entire with thin stalks inclined outwards and narrowed subapically ( Fig. 245 View Figs 245–265 ).
SPINNERETS. PMS: length 0.71, diameter 0.34. PLS: maximal diameter 0.80; length of basal, medial and apical segments 1.47, 0.73, 0.80; total length 3.00; apical segment shortly digitiform.
The length of the carapace varies from 5.80 to 6.25 in males and from 7.73 to 8.39 in females. The colouration varies through specimens very narrowly. The number of maxillary cuspules ranges from 32 to 40 in males and from 65 to 70 in females. The variation in the shape of the palpal organ and the length of the embolus seems to be almost imponderable (see Figs 206–207 View Figs 206–214 ).
The species occurs in the piedmont sandy desert (Tigrovaya Balka Nature Reserve) and in the woodless steppe foothills near Ganjina. Curiously, despite intensive collecting, A. castanea sp. nov. has never been found in the less harsh montane zone of the Aruktau Mts, occupied by A. incana and some other congeners.
The species is known from far southwestern Tajikistan only ( Fig. 371 View Fig ).
Andreeva (1968) included both conspecific females in the type series of Brachythele karatauvi . To avoid a situation of the same specimen being listed as a paratype in two different species, these females are not included in the types of Anemesia castanea sp. nov., despite definitely belonging to the latter congener.
Departamento de Geologia, Universidad de Chile
Department of Paleozoology, Swedish Museum of Natural History
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