Campsicnemus limnobates Evenhuis

Evenhuis, Neal L., 2009, Review of Campsicnemus (Diptera: Dolichopodidae) of the Marquesas, French Polynesia, with description of four new species groups, Zootaxa 2004, pp. 25-48 : 30-31

Campsicnemus limnobates Evenhuis

( Fig. 4 )

Campsicnemus limnobates Evenhuis, 2000: 293 . Yang et al. 2006: 461 [incorrectly giving the type locality as “ USA: Hawaiian Islands”].

Diagnosis. This species is easily distinguished from the others in the gladiator group by the mid and hind femora possessing long setae and cilia on their apical halves; and IIt1 possesses a single strong spatulate seta apically. It is separated from C. scurra and C. uncleremus by the yellow on the lower portion of the anepisternum (all brown in C. scurra and C. uncleremus ).

Redescription. Male. Body length: 2.15–2.62 mm. Wing length: 2.46–2.89 mm.

Head. Black, face dark brown to black with reddish highlights, paler brown near clypeus; oc and vt black, about two-thirds length of antennal arista; front, occiput, and vertex black with blue highlights; face constricted at middle, almost holoptic, eyes separated below antennae by width of 1–2 ommatidia; palp small, dark brown; proboscis brown, extending below eye in lateral view; antennal segments black to brown; postpedicel long, lanceolate, length about 3.5 x width; arista slightly longer than head height.

Thorax: Mesoscutum, scutellum, dark brown, with a few metallic greenish highlights laterally; pleura dark brown to brown except for yellow on the following sclerites: anepisternum, katepisternum, meron, and anepimeron; katetergite black; thoracic setae black: 4 dc; 2 np; 2 ph; 1 pa; 1 sc; ac absent; halter stem white, knob dark brownish black.

Legs: CI yellowish white, with apical tuft of three strong black setae, approximate basally, almost fused apically so as to appear as a single strong seta; CII mostly brown, some yellow anteriorly; CIII brown, slightly paler than surrounding pleura; FI and FII yellowish brown, FIII yellowish brown basally, becoming darker brown on apical one-fourth; remainder of legs brown; TiI yellow ventrally, brown dorsally, with long black seta subapically on ventral surface (MSSC); FI and remainder of foreleg unmodified, without MSSC; FII slightly bowed with medium sized setae basally, with subapical brown patch harboring two small setae; FIII slightly bowed with swollen basal area containing 2 small black setae and 1 long setae, middle of FIII swollen, containing very long curved (basalmost), wrinkled (medial), and straight (apicalmost) setae on ventral surface (MSSC); TiII slightly bowed and greatly swollen just beyond middle, with long hairs medially on dorsal surface, short hairs along ventral surface (MSSC). TiIII unmodified, without MSSC. IIt1 yellowish basally, blackish apically, short, about one-third length of IIt2, with strong black, spatulate apical seta and single associated strong black seta (MSSC). Remainder of leg segments without MSSC.

Wing: pale smoky throughout.

Abdomen. Dark brown with short black hairs dorsally on each tergite, a few longer hairs laterally; tergal interstices whitish; sternites yellowish brown. Hypopygium brown with paler brown cerci, not dissected.

Female. As in male except for lack of MSSC; postpedicel length about 1.5 x width; legs normal, without modifications; FI and FII brown.

Types. Holotype ɗ, FRENCH POLYNESIA: Marquesas Is: UA HUKA: Vaiee Cascade on Vaipaee River, 700 ft [213 m], water temp. 22.5 °C, 8°53'52"S, 139°33'17"W, skating on pools, 3 Nov 1999, D.A. Polhemus (USNM). Paratypes: FRENCH POLYNESIA: Marquesas Is: UA HUKA: 1 ɗ, 5 Ψ, same data as holotype (BPBM, USNM); 6 ɗ, 2 Ψ, Vaihonu Cascade and rocky streams in upper Hane Valley, 500 ft [152 m], water temp. 23 °C, 8°54'50"S, 139°32'15"W, 1 Nov 1999, skating on pools, D.A. Polhemus (BPBM, USNM).

Discussion. This species is a water skater found on pools and streams in the southern portion of the island of Ua Huka at elevations from ca. 150–215 m [500–700 ft].

Figures

FIGURES 3 – 6. Campsicnemus gladiator group, male mid basitarsi. 3. C. gladiator, n. sp. 4. C. limnobates Evenhuis. 5. C. scurra Parent. 6. C. uncleremus Evenhuis.