Campsicnemus englundi Evenhuis

Evenhuis, Neal L., 2009, Review of Campsicnemus (Diptera: Dolichopodidae) of the Marquesas, French Polynesia, with description of four new species groups, Zootaxa 2004, pp. 25-48 : 37-38

Campsicnemus englundi Evenhuis , new species

( Figs. 10 , 15 )

Diagnosis. Most similar in appearance to C. plautus , n. sp. but can be easily distinguished by the mid basitarsus thick but not flared apically (mid basitarsus flared in C. plautus ) and by the scattered strong setae on the mid tibia (setae of uniform length on the anterior surface and longer and denser on the basal third of the posterior surface in C. plautus ).

Description. Male. Body length: 2.2–2.4 mm. Wing length: 2.1–2.6 mm.

Head. Black, face dark brown to black with reddish highlights, paler brown near clypeus; oc and vt black, about one-third length of antennal arista; front, occiput, and vertex black with blue highlights; face constricted at middle, almost holoptic, eyes separated below antennae by width of 1–2 ommatidia; palp small, dark brown; proboscis brown, extending below eye in lateral view; antennal segments black to brown; postpedicel long, lanceolate-conical, length about 3 x width; arista slightly longer than head height.

Thorax: Mesoscutum, scutellum, and pleura dark brown to black throughout, with a few metallic bluish highlights dorsolaterally; thoracic setae black: 1 + 4 dc; 2 np; 2 ph; 1 pa; 1 + 1 sc; ac absent; halter stem white, knob dark brown.

Legs: CI and FI yellowish; remainder of legs brown; foreleg unmodified, without MSSC; FII ( Fig. 15 ) with strong, swollen area ventrally bearing 4–5 strong, stiff black setae ventrally, patch of brown hairs basally; with prominent protuberance on apical one-third bearing small patch of short slightly curved black setae; FIII as in C. gladiator , with 7–8 long setae on ventroapical surface (MSSC); TiII ( Fig. 15 ) thin basally, slightly swollen on apical two-thirds, with 6 strong black setae on apical third on dorsal surface, small patch of stiff black setae on mesal surface at point where thickness of tibia starts (MSSC), a few smaller fine black setae and hairs along medial third of mesal surface. TiIII with row of long stiff setae on basal half, becoming shorter apically, dorsal surface with small hairs and only two long setae (one subbasally; the other near middle) (MSSC). IIt1 ( Fig. 14 ) short, broad, shield-like apically, about one-half length of IIt2; IIt2 inserted apically on IIt1. Remainder of leg segments without MSSC.

Wing: pale smoky throughout; posterior crossvein perpendicular to vein M.

Abdomen. Brown, darker brown on posterior margins, with short black hairs dorsally on each tergite; tergal interstices whitish; sternites pale brown. Hypopygium brown with paler brown cerci, not dissected.

Female. Unknown.

Types. Holotype ɗ (BPBM Type 16,641) [BPBM1013180] and 3ɗ [BPBM1013181] paratypes (one headless) from FRENCH POLYNESIA: Marquesas: FATU HIVA: North Branch Omoa River, 400–500 ft [122–152 m], 26 Aug 2001, riffles, pools, R.A. Englund (see Fig. 10 ). Other paratype: FRENCH POLYNESIA: Marquesas: FATU HIVA: 1ɗ [BPBM1013182], Omoa River, 1000–1320 ft [328–402 m], 28 Aug 2001, cascade/pool, R.A. Englund; 1ɗ, Upper Omoa River, 1.6 mi [1 km] above Omoa, 10°30'31"S, 138°39'46"W, 26 Aug 2008, skating on surface of stream pools, D.A. Polhemus. Holotype and paratypes in BPBM.

Etymology. Named in honor of friend and colleague, Ronald Englund, for his successful efforts at collecting new and unusual riparian dolichopodids throughout French Polynesia over the last couple of decades.

Figures

FIGURES 10 – 13. Marquesan habitats. 10. Fatu Hiva, Omoa River: type locality for Campsicnemus cheesmanae, n. sp. and C. englundi, n. sp. 11. Ua Pou, Pouaoa River: habitat for C. gladiator, n. sp. 12. Hiva Oa, Vaiao River: habitat for C. uncleremus Evenhuis and C. dytei, n. sp. 13. Hiva Oa, Mt. Temetiu: habitat for C. hihiroa, n. sp. and C. taratara, n. sp.

FIGURES 14 – 16. Campsicnemus englundi group, male midlegs. 14. C. cheesmanae, n. sp., femur, tibia, and basitarsus. 15. Campsicnemus englundi, n. sp., femur, tibia, and basitarsus. 16. Campsicnemus plautus, n. sp., femur, tibia, and all tarsi, showing flared basitarsus and swollen second tarsomere.