Tokudaia Kuroda, 1943

Guy G. Musser & Michael D. Carleton, 1993, Order Rodentia - Family Muridae, Mammal Species of the World (2 nd Edition), Washington and London: Smithsonian Institution Press, pp. 501-755 : 670

Tokudaia Kuroda, 1943 . Biogeographica, 139:61.

TYPE SPECIES: Rattus jerdoni osimensis Abe, 1934 .

SYNONYMS: Acanthomys ( Tokuda, 1941, not Lesson, 1842), Tokudamys .

COMMENTS: In addition to two species listed below, a third, yet to be named and described, occurs on Tokun-oshima Isl in the Ryukyu group ( Tsuchiya, 1981; Tsuchiya et al., 1989). Best estimate of phylogenetic relationships, based on dental morphology, placed Tokudaia in a group containing Apodemus , Pliocene Rhagapodemus , and Quaternary Rhagamys ; and suggested that Tokudaia evolved from Miocene Parapodemus-Apodemus ancestral stock; its evolutionary link to primitive species of Apodemus may be found in Miocene or Pliocene sediments of China (see Kawamura, 1989).