Maxomys whiteheadi (Thomas, 1894)

Guy G. Musser & Michael D. Carleton, 1993, Order Rodentia - Family Muridae, Mammal Species of the World (2 nd Edition), Washington and London: Smithsonian Institution Press, pp. 501-755 : 614

Maxomys whiteheadi (Thomas, 1894) . Ann. Mag. Nat. Hist., ser. 6, 14:452.

TYPE LOCALITY: Malaysia, Sabah (N Borneo), Gunung Kinabalu .

DISTRIBUTION: Peninsular Thailand south of Isthmus of Kra, Malay Peninsula, Sumatra, Borneo, and various adjacent islands; absent from Java and Bali.

SYNONYMS: asper , batamanus , batus , coritzae, klossi , mandus , melanurus , melinogaster , perlutus, piratae, subitus.

COMMENTS: A Sundaic endemic. Ellerman and Morrison-Scott (1951) included whiteheadi in the Sulawesi musschenbroekii , but the two are separate species ( Chasen, 1940; Medway, 1977; Musser, 1991; Musser et al., 1979; Tate, 1936). Spermatozoal morphology ( Breed and Yong, 1986), and data from biochemical, morphological, and cytological studies ( Chan et al., 1978, 1979; Yong, 1969) pointed to a close relationship between M. whiteheadi and the Malayan M. inas .