Leptopsilopa pollinosa (Kertész), Kertesz

Zatwarnicki, Tadeusz, Cielniak, Magdalena & Pochrząst, Katarzyna, 2016, A revision of the Old World species of Leptopsilopa Cresson (Insecta: Diptera: Ephydridae), Zootaxa 4093 (3), pp. 343-362 : 354-360

Leptopsilopa pollinosa (Kertész)

( Figs. 7 B–10)

Ephygrobia pollinosa Kertész, 1901: 81 .

Psilopa pollinosa: Hendel, 1913: 97 .

Leptopsilopa pollinosa: Cresson, 1925: 251 .

Psilopa irregularis Malloch, 1934: 314 —synonymy by Cresson, 1936: 53. Psilopa confentei Canzoneri and Munari, 1991: 163 , new synonym. Psilopa confenteae: Mathis and Zatwarnicki, 1995: 40 [correction; name formed to honor a woman]

Description. Body length 1.58–1.81 mm. Head ( Fig. 9 A–9B). Antennae dark, basiflagellomere lighter in ventral half; arista with seven-nine dorsal rays; frons shining blue, with more golden reflection medially, frontal setae equal in length, slightly shorter than ocellar; face narrow, its width is equal to distance between antennae (facial ratio 0.11), wrinkled transversally and shiny medially, silvery microtomentose laterally; facial seta moderate, in lower 1/3 of face, parafacialia whitish microtomentose; palpus black; gena-to-eye ratio 0.05–0.08.

Thorax. Mesonotum and scutellum black shiny, anepisternum and sides of mesonotum with golden reflection, remaining with black reflection; scutellum trapezoidal with rounded posterior margin, surface flat, bears a few short setulae; wing hyaline; costal vein ratio 0.87–1.18; M vein ratio 0.49–0.61; legs black, mid and hind tarsi yellowish, except dark distal segment; stem of halter brown, head white.

Abdomen black, metallic shiny with green and golden reflection. Male terminalia ( Figs. 8 A–8D). Epandrium in ventral view ( Fig. 8 A) U-shaped with dorsal arch slightly narrower than lateral arms; in lateral view ( Fig. 8 B) height almost 2x the greatest width, pouch-like; cercus in ventral view ( Fig. 8 A) uniformly oval; in lateral view ( Fig. 8 B) shallowly arched; pregonostylus well-developed, in ventral view ( Fig. 8 A) spatulate with bar-like, lateromedial basal projection bearing 4 long and 3 much smaller setulae; in lateral view ( Fig. 8 B) C-shaped, with truncate base and rounded apex, ventral margin almost straight, dorsal margin deeply concave, projection almost oval; postgonostylus in ventral view ( Figs. 8 C) elongate, inverted L-shaped, bearing setulae on apical half; in lateral view ( Figs. 8 D) finger-like and slightly arcuate; medandrium (= subepandrial plate) ( Figs. 8 C–8D) bandlike, arch-like; aedeagus in ventral view ( Figs. 8 C) broadly elliptical, tapered gradually to apex, small indentation on 2/3 length; in lateral view ( Figs. 8 D) elongate, funnel-like, wider basally, tapered to rounded apex; gonite bears three well developed setae, in ventral view ( Fig. 8 C) spatulate with sinuous lateral margin; in lateral view ( Figs. 8 D) narrowly ovate; phallapodeme in ventral view ( Fig. 8 C) goblet-shaped; in lateral view ( Figs. 8 D) lunate with small subrectangular keel apically; hypandrium in ventral view ( Figs. 8 C) trapezoidal, with convex posterior margin; in lateral view ( Fig. 8 D) subtriangular, narrowing toward phallapodeme.

Type material examined. Lectotype female of senior synonym, herein designated, labelled: " Singapore Biró 1898", " Ephygrobia pollinosa Kert. det Kertesz typus " (red), " typus " (with red margin) in good condition and a paralectotype female in good condition, labelled: " Singapore Biró 1898", " Ephygrobia pollinosa Kert. " "det Kertesz typus " (red), " typus " (with red margin) preserved in HNHM; paratype female of Malloch's species in moderate condition (abdomen flattened laterally and left wing missing) labelled “Para- type ” (round label with yellow margin), “W. M. Gifford Collection” (red printed letters), “Pres. by P. A. Buxton B. M. 1934-97 ”, “ Psilopa irregularis Malloch Paratype ” (handwritten), “H. C. Kellers Coll. IV-18 ” (IV-18 handwritten), “ 760–900 ft. Tutuna Samoa ” (numbers handwritten) preserved in BMNH; holotype of L. confentae labelled: " NEPAL Bagmati River near Kathmandu 1300 m, 4. VI 1980 A.M.CONFENTE L.GARIGGIO legit" (with black margin), " HOLOTYPUS Psilopa confentei ♂ sp. nov. S. Canzoneri - L. Munari" (red with black margin, handwritten, after HOLOTYPUS black line), "MUSEO CIVICO DI STORIA NATURALE DI VENEZIA Materiale tipico descritto Inv. n. 02933" (orange, number handwritten) and a paratype male with the same locality label preserved in MCSNV.

Other material examined: AMERICAN SAMOA: Pago Pago, Tutulla, VII.–VIII. 1962, M. R. Wheeler, 1 ♂, III.–IV. 1962, M. R. Wheeler, W. S. Stone, 2 ♂, 2 ♀, and I. 1980, N. L. H. Krauss, 1 ♀. AUSTRALIA: New South Wales: Mosman, stream margin, 21. IV. 1957, W. W. Wirth, 1 ♂, 1 ♀; Red Rock, Corindi Cr., coast swamp forest, 7. XI. 1980, D. J. Bickel, 2 ♂, 1 ♀ (all USNM). Northern Territory: Katharine, at light, 17. VIII. 1973, L. P. Kelsey, 5 ♂, 8 ♀ (ANIC); 22 km W of Humpty Doo, 10. VI. 1964, K.R. Norris, 1 ♂; Lee Point, VI. 1964, K.R. Norris, 1 ♀; Fogg Dam area, 9. IV. 1980, G. F. Hevel, J. A. Fortin, 1 ♂, 8 ♀ (USNM). Queensland: 6 km N on Emerald Ck. Rd., S of Mareeba, 490 m, 25. IV. 1990, J. Heraty, 1 ♀ (CNC); Annan R., 3 km W by S of Black Mt., at light, 26. IV. 1981, D. H. Colless, 1 ♀; Hann R., 15.12S, 143.52E, at light, 27. VI. 1986, D. H. Colless, 1 ♀; Darr River, 31 km NW by N of Longreach, at light, 7. IV. 1976, D. H. Colless, 1 ♀; Laceys Creek, Mission Beach, 17.54S, 146.06E, 13–14. V. 1980, I. D. Naumann, J. C. Cardale, 1 ♂; Shiptons Flat, 15.47S, 145.14E, 16. V. 1981, D. H. Colless, 1 ♀; 3.5 km SW by S of Mt. Balrd, 15.10S, 145.07E, light trap, 3. V. 1981, D. H. Colless, 1 ♀; 4 km SSE of Cape Tribulation nr. Daintree, at light, 21–22. XI. 1981, D. H. Colless, 1 ♂, 1 ♀; Cooktown, 1 km W, 15.28S, 145.15E, 12. V. 1981, D. H. Colless, 1 ♂; 11 km ENE of Mt Tozer, 12.43S, 143.18E, at light, 11–16. VII. 1986, D. H. Colless, 1 ♀; 1 km N of Rounded Hill, 15.17S, 145.13E, at light, 5–6. V. 1981, D. H. Colless, 2 ♀; 2 km S of Horseshoe, L/out Blackdown Tableland, 23–24. IV. 1981, D. H. Colless, 1 ♀; 3 km NE of Mt Weeb, 15.03S, 145.09E, Malaise trap, 30. IV.–2. V. 1981, D. H. Colless, 4 ♀; 3 km NE by N of Julatten, 16.35S, 145.22E, at light, 26. IX. 1980, D. H. Colless, 2 ♀ (all ANIC); Cairns, 19–22. IV. 1957, W. W. Wirth, 4 ♂, 3 ♀ and 6. X. 2002, A. Freidberg, 3 ♂, 3 ♀; Curtain Fig Tree, Rt. 52, 2 km S Yungaburra, 11. X. 2002, A. Freidberg, 6 ♂, 4 ♀; Fig Tree Creek, 39 km S Ingham, 14. IV. 1980, G. F. Hevel, J. A. Fortin, 2 ♀; 7–14 km W of Herberton, via Watsonville, 1. V. 1967, D. H. Campbell, 1 ♂; Maitland Downs, 3 km S, 15. IV. 1980, G. F. Hevel, J. A. Fortin, 1 ♂; Bell Bird Cr., Lamington Nat. Park, 30. V. 1966, Z. Liepa, 1 ♂; Brisbane, 30. XI. 1900, L. Biró, 1 ♂, 1 ♀ (all USNM); 7. X. 1964, G. B. Monteith, 1 ♂, 5 ♀ (CNC); Deeral, Rt. 1, 4 km S Cairns, 10. X. 2002, A. Freidberg, 1 ♂; Josephine Falls, 14. IV. 1980, G. F. Hevel, J. A. Fortin, 1 ♂, 1 ♀; Kuranda, 6. X. 2002, A. Freidberg, 6 ♂, 7 ♀; 1 km E Mt. Molloy, 15. IV. 1980, G. F. Hevel, J. A. Fortin, 2 ♂; 3 km Woodstock, 14. IV. 1980, G. F. Hevel, J. A. Fortin, 1 ♂, 1 ♀ (all USNM); Iron Range, 12–14. VI. 1971, J. Feehan, 1 ♂, 4 ♀; Crystal Creek, 43.4 km SWS of Ingham, 8. VII. 1971, Z. Liepa, 1 ♀ (ANIC); Woombah, 200 km S of Brisbaine, 22–30. IX. 2002, M. Bartak, 1 ♂, 1 ♀ (CULSP); Wongabei Forest, Atherton, 19. IX. 2002, Š. Kubik, 1 ♂, 1 ♀ (CULSP). Tasmania: Mt. Windsor T'land, 1250 m, 17. IV. 1994, B. J. Sinclair, 1 ♀ (CNC). Victoria: Broomfield, 2. XI. 1972, J. Sedlaczek, 2 ♂, 1 ♀ (CNC). Western Australia: Cape Bertholet, 8 km S, West Kimberley, 19. IV. 1977, D. H. Colless, 1 ♂ (ANIC). BRUNEI: Bandar Seri Begawan, prairie, 13. IV. 1993, P. Grootaert, 1 ♂, 1 ♀ (RBINS). BURMA (= Myanmar): Shingbwiyang, 29. III. 1944, L. C. Kuitert, 2 ♂, 2 ♀ (USNM); Ze Yar, 1300 m, forest cleaning, 7–8. II. 1998, H. Nickel, 1 ♂, 1 ♀ (HNHM). CAMBODIA: Phnom Penh, garden, MT+PT, 11º30'34”N, 104º53'54”E, 31.VII.– 15.VIII. 2013, M. Petryl, 12 ♂, 2 ♀ (CULSP); Siem Reap, XI. 1958, N.L.H. Krauss, 2 ♂, 1 ♀ (BMNH). CHINA: Fukien, Chung An, Bohea Hills, 24. VIII. 1960, T.C. Maa, 1 ♂, 1 ♀, 10. XI. 1939, 1 ♂, 2 ♀ (BBM). FIJI: Lautoka, 13. IX. 1919, W. Greenwood, 1 ♀ (all BMNH); Navai-Eteani (Naitrasin Prov.), 734 m, 7. V. 2003, 3 ♀ (CNC); Navua, VI., 1926, L. Stoner, 2 ♀; Suva, VI. 1963, M. R. Wheeler, 1 ♀ (USNM), 1 ♀ (BMNH); Viti Levu Nandi, 0– 100 m, 5. III.–IV. 1974, N.L.H. Krauss, 14 ♂, 9 ♀ (USNM). GUAM: Agana, 23. V. 1948, K. L. Maehler, 1 ♀ and X. 1952, N.L.H. Krauss, 1 ♀; Anderson AFB, X. 1957, N.L.H. Krauss, 2 ♂, 1 ♀; Matizo, X. 1957, N. Krauss, 4 ♂, 1 ♀; Nimitz Beach, VIII. 1952, N.L.H. Krauss, 1 ♀; Oca Point, 2. VI. 1945, H. S. Dybas, 1 ♂; Talofofo, II. 1958, N. Krauss, 2 ♂; Umatac, X. 1957, N. Krauss, 4 ♂, 2 ♀; Ylig Bay, XII. 1958, N.L.H. Krauss, 2 ♂; Yigo, X. 1957, N. Krauss, 2 ♂, 1 ♀ (all USNM), 1 ♀ (BMNH). INDIA: Andhra Pradesh: Hyderabad, 26. X.–5. XI. 1971, A. C. Pont, 14 ♂, 8 ♀ (BMNH); Assam: 10–16 km N Tinsukia, 6.–29. III. 1944, D. E. Hardy, 10 ♂, 8 ♀; Chabua, 9. I.– 20. II. 1944, D. E. Hardy, 5 ♂, 1 ♀; Deamoolie, 16 km W Doom Doom, 20. V. 1944, D. E. Hardy, 3 ♂, 6 ♀; Rupsi, 23 km NW Dhubri, 3. XI. 1943, D. E. Hardy, 3 ♂; Sadiya, 13. II. 1944, D. E. Hardy, 5 ♂, 1 ♀; Bengal: Calcutta, 14–15. IV. 1980, A. Freidberg, 6 ♂, 12 ♀; Calcutta, X. 1958, N.L.H. Krauss, 2 ♂, 1 ♀; Calcuta, Eden garden, 22– 25. XII. 1956, A. P. Kapur, 1 ♂, 2 ♀; Karnataka: Bangalore, 916 m, host: Laphygma ( Lepidoptera : Noctuidae ), II. 1978, Veeysch, 1 ♀; Bangalore, 3. IV. 1980, W. N. Mathis, A. Freidberg, 7 ♂, 7 ♀;: Bangalore, University Campus, 12º8''N, 77º37'E, 1.–15.II. 2005, J. Olejniček, 1 ♂, 2 ♀ (CULSP); Mudigere, 6–7. IV. 1980, W. N. Mathis, A. Freidberg, 5 ♀; Kerala: 30 km, SE Munnar Lake, 1300 m, 27. XI. 2002, A. Freidberg, 9 ♂, 7 ♀ (all USNM); Maharashtra: Mumbai (“ Bombay ”), 1902, L. Biró, 1 ♂ (ANSP); Tamil Nadu: Tondiarpet, Madras, over open sewage, VIII. 1913, Fletcher, 1 ♂; Yercaud, nr. Salem, 1370 m, 8.–10. IV. 1962, G.J. Spencer, 2 ♀ (all BMNH); Uttar Pradesh: Agra, Shah Jehan Gardens, swept on shores on small ponds and a watered grassy area, 25. XI. 1989, L. Papp, 1 ♂; Keetham Lake, 20 km N of Agra, swept on shore-lake mud, 27. XI. 1989, L. Papp, 1 ♂ (HNHM). INDONESIA: Flores: Bajawa, 1200 m, 17. VI. 1927, Sunda Exp. Rensch, 18 ex.; Rana Mese, 20–30. VI. 1927, Sunda Exp. Rensch, 1 ♀ (SDEI); Java: Jakarta, light, III.–VI. 1955, O. M. Recs, 1 ♀ (USNM); Lombok: Ekas, IV. 1927, Sunda Exp. Rensch, 1 f; Selóng, 21. IV. 1927, Sunda Exp. Rensch, 3 ♂; Sumbawa: Dompu, 24–25. V. 1925, Sunda Exp. Rensch, 38 ex.; Sumbawa-Besar, 24. IV.–2. V. 1927, Sunda Exp. Rensch, 1 m; Wawu, 450 m, IV. 1927, Sunda Exp. Rensch, 1 ♂, 1 ♀ (all SDEI). LAOS: Vientiane, at light, 27. V. 1965, P.D. Ashlock, 2 ♂, 4 ♀, 8. V. 1965, 1 ♂, Ban Theuong, 18 km NW of Xieng Khonang, 1035 m. 2–6. VIII. 1960, at light, R.E. Leech, 1 ♀, Muong Sing, NW of Luang Prabang, 650 m., 6–10. 1960, S. & L. Quate, 7 ex.; Pakse (Sedone Prov.), 11–22. V. 1965, P. D. Ashlock, 2 ♂, 1 ♀. MALAYSIA: Sarawak, Kuching, Santubong, 797–1500 m, Dancing in air over Intertidal zone at dusk, 26. VI. 1958, T. C. Maa, 1 ♂ (all BBM); Sabah: Tanjung Aru Beach, 3–20. VIII. 1983, G.F. Hevel, W. E. Steiner, 2 ♂, 5 ♀. MICRONESIA: Yap: Map I., 22. X. 1952, N.L.H. Krauss, 2 ♂, 1 ♀; Yap I., Colonia, 23. IV. 1954, J. W. Beardsley, 2 ♂, 1 ♀, on grass, 1. III. 1949, K.L. Maehler, 1 ♂; Yap I., IX. 1952, N.L.H. Krauss, 12 ♂, 9 ♀ (all USNM) and 1 ♂, 1 ♀ (BMNH); Yap I., Gagil Dist, VII.–VIII. 1950, R.J. Goss, 1 ♀; Yap I., Giliman, 11. VI. 1957, C. W. Sabrosky, 3 ♂, 1 ♀; Yap I., Ruul, nr. Colonia, 13. VI. 1957, C. W. Sabrosky, 1 ♂. NEPAL: Chomrung (3 km W), Kyumnu Khola (Kaski Dist.), 1800 m, 26. X. 1985, W.N. Mathis, 1 ♀; Nagarkort, 2200 m, 2. V. 1980, A. Freidberg, 1 ♀; Pokhara, 900 m, 3–4. XI. 1977, G. F. Hevel, 4 ♀ (all USNM); Pokhara, 19 km NW, 2000 m, grassy meadow. 7. IV. 1983, U. Gärdenfors, 2 ♀; Royal Chitwan Nat. Park, 2 km NW Elephant Camp, grassy meadow, 2. IV. 1985, R. Hall, 37 ex.; Royal Chitwan Nat. Park, Eastern Park Jungle Lodge Sal Forest near Lodge, 3. IV. 1985, R. Hall, 17 ex. (all MZLU); Royal Chitwan Nat. Park, 250 m, 1–2. XI. 1977, G. F. Hevel, 4 ♀; Sauraba (Chitwan Dist.), 2 km E Dhungar Khola, grassy bank, 31.X.–1. XI. 1985, W. N. Mathis, 1 ♂, 8 ♀ (all USNM); Godavari (near Kathmandu), 1525–1830 m, 15. IV.–15. VIII. 1967, Can. Nepal Exp., 15 ♂, 19 ♀; nr Birganj Lothar, 1370 m, 10. IX. 1967, Can. Nepal Exp., 1 ♀ (all BMNH). NEW CALEDONIA: Aoupinié, 21°08.941’S 165°19.407’E, refuge 420 m rainforest, at light, 7–29. III. 2008, T. Blaik, R. Dobosz, 18 ♂, 16 ♀; Bois du Sud, 22°10.443’S 166°45.760’E, camp 210 m, riverside, rainforest margin, at light, 3–9. III. 2008, T. Blaik, R. Dobosz, 3 ♂, 1 ♀; Col d’Amieu, 21°35.407’S 165°47.728’E, 420 m, rainforest/planted forest, at light, 15–16. III. 2008, T. Blaik, R. Dobosz, 1 ♂, 2 ♀; Farino, 21°38.533’S 165°46.477’E, refuge 300 m, garden vegetation/forest, at light, 10. III. 2008, T. Blaik. R. Dobosz, 1 ♂; Parc des Grandes Fougères Farino env. 21°37.632’S 165°45.830’E, 467 m, rainforest, at light, 13. III. 2008, T. Blaik, R. Dobosz, 1 ♂, 1 ♀; Tchamba (Wâo Uni), 21°00.332’S 165°14.911’E, refuge 396 m, rainforest, at light, 31. III. 2008, T. Blaik. R. Dobosz, 1 ♂, 1 ♀ (all TZPC); Mt. Koghis, 800 m., 1–6. IX. 1972, J. F. McAlpine, 10 ♂, 3 ♀; Col de Roussettes, 23 km N Bourail, 350 m, ex dung trap, 17. VII. 1995, B. J. Sinclair, 1 ♂; Ouenguip, 15 km W Hienghene, 17. VII. 1995, B. J. Sinclair, 1 ♀ (CNC). NORTHERN MARIANA ISLANDS: Tinlan I., II. 1958, N.L.H. Krauss, 4 ♂, 1 ♀ and 15 ex. in plastic container (USNM); 11. XI. 1952, J. W. Beerdsley, 1 ♀; Saipan, Med. Bn 2nd Mar. Div., 6. IX. 1944, 2 ♂, 1 ♀ (all BMNH). OMAN: Al Kabura, 10. IV. 1985, P. Ardö, 3 ♂, 6 ♀; Barka beach, 8. IV. 1985, P. Ardö, 8 ex.; Ghuzayn, date cult, 10. IV. 1985, P. Ardö, 1 ♀; Mazara, 6. IV. 1985, P. Ardö, 1 ♂; Muscat, 10. IV. 1985, P. Ardö, 59 ex.; Qurm, 5–9. IV. 1985, P. Ardö, hotel garden or wadi beach, 17 ex. (all MZLU); Dhofar, Khor Rawri, on Scirpus marina (Cyperaceae) , 12. XI. 1992, J. C. Deeming, 1 ♂ (NMWC); Dhofar, Ain Hamran, 10. X. 1990, M.J. Ebejer, 1 ex; Sharqiya, Wadi Bani Khalid, 24. II. 1990, M.J. Ebejer, 9 ex. (MJE). PALAU ISLANDS: Babelthuap I., Ngardmau, 10. V. 1957, C. W. Sabrosky, 1 ♀; Babelthuap I., Ngaremlangui, 3. VI. 1957, C.W. Sabrosky, 1 ♂; Babelthuap I., Ulimang, sweeping roadside vegetation, 19. XII. 1947, H.S. Dybas, 1 ♀; Islet near Koror I., IX. 1952, N.L.H. Krauss, 2 ♀; Koror I., 13. IV.–3. V. 1957, C. W. Sabrosky, 11 ♂, 20 ♀ (all USNM), 2 ♂ (BMNH); Malakal I., on wet rocks, 2. V. 1957, C. W. Sabrosky, 1 ♂, 3 ♀; Ngesebus I., 29. V. 1957, C.W. Sabrosky, 2 ♂, 1 ♀. PAPUA NEW GUINEA: Central Papua Prov.: Kupiano, grass, 25. VII. 1982, J.W. Ismay, 1 ♂, 3 ♀ (all USNM); Laloki, under banana, 20. VII. 1982, J.W. Ismay, 1 ♀ (BMNH); Latoki, low vegetation, 13. VII. 1982, J. W. Ismay, 2 ♂ (USNM); Manus (Bismarck Isl.): Lorengau, 24. VI. 1962, Noona Dan Exp. 61–62, 1 ♀ (ZMUC); Morobe Prov.: Bubia, marsh, 30. IV. 1986, J. W. Ismay, 3 ♂, 2 ♀ (USNM); Herzog Mts, Vagau, 1220 m, 4.–17. I. 1965, 5 ♂, 4 ♀ (all BMNH); Nat'l Capital Dist.: Konedobu, under trees, 15. V. 1981, J.W. Ismay, 1 ♀; Saraga, garden, 24. II. 1985, J.W. Ismay, 1 ♂, 1 ♀; Oro Prov.: Nr. Tufi, grassland, 3. VIII. 1983, J.W. Ismay, 3 ♂, 3 ♀ (all USNM); Western Highlands Prov.: Baiyer River, rain forest, 1220 m, 2–5. XI. 1973, K.A. Spencer, 1 ♀ (BMNH); Mt. Hagen, 1700 m, 1973, G. Baker, 2 ♀ (CNC). PHILIPPINES: Palawan, Puerto, Princessa, XI. 1925, R.C. McGregor, 1 ♀ (BMNH). SAMOA: 1 ♂ (NMW), Vailina, 19. XI. 1950, R. Harrison, 1 ♀ (USNM); Upolu, Apia, 0–150 m, IV. 1977, N.L.H. Krauss, 1 ♂. SOLOMON IS.: Guadalcanal, Honiara, XII. 1980.– I. 1981, N.L. H. Krauss, 1 ♂; New Georgia, Munda, XI. 1980, N.L.H. Krauss, 1 ♂ (all BMNH); 0–200 m, I. 1974, N.L.H. Krauss, 4 ♂; Tulagi, 0–10 m, 11. XII. 1973, N.L.H. Krauss, 5 ♂ (all USNM); Rennell, Niupani, 17. VIII. 1962, Noona Dan Exp., 8 ♂, 5 ♀ (ZMUC). SRI LANKA: Eastern Prov.: Kokagala Mnt. 20 mls N Bibile, swept on dry grassland, stream, 13. III. 1962, P. Brinck, H. Andersson, L. Cederholm, 3 ♀; stream 15 mls SSW Batticaloa, dry meadow, 8. III. 1962, 1 ♂; E. Prov., Gal. Oya, alt. 90 m, 14 mls E Bibile, sandy grass, river, 8. III. 1962, 3 ♀; Vayiriuttu, 5 mls W Trincomallee, grassy ground, 9. II. 1962, 1 ♂, 1 ♀; Northern Prov.: 7 mls E Mankulam, swampy tank, swept on veg. in reservoir (tank), 14. II. 1962, 2 ♀; E Mankulam, small stream, swept on muddy ground at pond, 14. II. 1962, 2 ♂, 2 ♀; 10 mls SE Mannar, gian's tank, swept on veg. in reservoir (tank), 15. II. 1962, 5 ex.; Kudattanal, 6 mls SE Point Pedro, swept on muddy ground at pond, 13. II. 1962, 1 ♀; 2 mls E Paralyanalankulam, 20 mls W Vavuniya, at pond in dry forest, 15. II. 1962, 2 ♂, 1 ♀; Per Aru, 9 mls E Mankulam, swept on grass in forest, 14. II. 1962, 1 ♂, 1 ♀; North Western Prov. Dedura Oya, 5 mls NE Kurunegala, river, swept above surface of stream, 7. II. 1962, 1 ♂; Swamps 10 mls E Puttalam, swept along roads in marshy land, 2. II. 1962, 1 ♀; 7 mls NE Puttalam, swept on dry grassland, 1. II. 1962, 1 ♀; North Central Prov.: 3 mls S Minneriya, swept in the veg. small stream, 11. II. 1962, 1 ♀; Wilpattu N. P. Maradan Maduwa, 23 mls W Anuradhapura, swept on moist grassland, 2. II. 1962, 5 ex.; stream at Maha, Bulankulama, 7 mls SW Anuradhapura, swept on moist grassland, 4. II. 1962, 6 ex.; Yan Oya, 24 mls W Trincomalee, swept on moist grassland, 10. II. 1962, 1 ♂, 2 ♀, swept on muddy ground along stream, 1 ♂; Centr. Prov.: Foothils of Knuckle Mnts,, 10 mls ENE Kandy, swept on veg. at streams in tea plantation, 11. III. 1962, 1 ♀; Rangala Knuckle Mnts, 12 mls ENE Kandy, swept along roads, 11. III. 1962, 1 ♀; stream 2 mls E Madugoda, 16 mls E Kandy, grassy ground, 12. III. 1962, 1 ♀; Maskellya Oya, 6 mls SW Hatton, swept along roads in tea plantation, 18. III. 1962, 6 ex.; 3 mls NW Hanguranketa, alt. 550 m, 10 mls SE Kandy, swept on grass coconut plantation, 15. III. 1962, 1 ♂, 1 ♀; Diyagama West at 1460 m, 8 mls S Nuwara-Ellya, swept on veg. at streams in tea plantation, 19. III. 1962, 8 ex. (all MZLU); Peradeniya Botanic. Gardens, 20. IV. 1980, W. N. Mathis, T. Wijesinhe, L. Jazawickrema, 2 ♂, 9 ♀ (USNM); Southern Prov.: Koskoda, 44 mls SSE Colombo, swept on herbs on sandy beach, 25. I. 1962, 1 ♀; Yoda Wawa at Tissamaharama, grassy ground, 22. III. 1962, 10 ex.; Sabaragamuwa Prov.: Hatherleigh, 1 mile S Rakwana, at 550 m, swept on grassland, 28. II. 1962, 1 ♂, 1 ♀ (all MZLU); Western Prov.: Bellanwila, 8. V. 1980, W. N. Mathis et al., 2 ♀; Colombo, 18. IV. 1980, W. N. Mathis, 1 ♀ (all USNM); Colombo, Colpetty, garden, 5–13. I. 1962, 6 ex.; Alawala, 26 mls NE Colombo, ravine with small stream, 17. I. 1962, 1 ♂, 2 ♀ (all MZLU); Henaratgoda, 14. III. 1902, H. Uzel, 1 ♂ (NMW); Nugegoda, 16. I. 1968, B. P. Karunaratne, 1 ♀ (BMNH); Labugama, 24 mls ESE Colombo, swept on grass in forest, 21. I. 1962, 2 ♀; Yakkala, 18 mls NE Colombo, swept along roads in paddy field distr., 15–31. I. 1962, 1 ♂, 1 ♀; Prov. of Uva: Diyatalawa, alt. 1220 m, 3 mls N Haputale, grassy ground, 21. III. 1962, 2 ♂; Mahavell Ganga at Alutnuwara, 24 mls E Kandy, swept on muddy ground at pond, 12. III. 1962, 1 ♀ (all MZLU). TAIWAN: Kwantzeling, Tainan Hsien, 250 m. 6–7. IV. 1965, C. M. Yoshimoto, 1 ♀ (BBM); Paroe, nordl. Paiwan-Distr., VIII. 1912, H. Sauter, 1 ♂, 1 ♀; Polisha, XII. 1908, H. Sauter, 1 ♂, 2 ♀ (all SDEI); Pilam, VIII. 1912, Sauter 1 ♂ (ANSP); Yang, Ming Shan, I. 1966, light, E. M. Bravi, 1 ♂ (BMNH). THAILAND: Chaiyaphum Prov.: Phukhieo Banlat, 16–20. XII. 1952, R. E. Elbel, 2 ♂; Chiang Mai: Chiang Mai, light trap, IV.–V. 1958, M. Notonanda, 1 ♀ (all USNM); Mae Taeng Elephant Camp, 50 km N Chiang Mai, on elephant dung, 29. X. 2004, L. Papp, M. Földvári, 1 ♂ (HNHM); Kanchanburi: Kanchanburi, 27. III. 1952, R. E. Elbel, 1 ♀; Krung Thep Mahanakhon (=Bangkok): Bangpo, light trap, VIII.–IX. 1962, J. Scanion, 1 ♀; Loei: NaPhung, Dansai, 10. VI. 1955, R. E. Elbel, 4 ♂, 5 ♀ (all USNM); Nan: Nan, 31. VIII. 1985, Reacher, Huber, 1 ♀ (CNC); Ratchaburi: Ratchaburi, 17. II. 1952, R. E. Elbel, 1 ♀; Songkhla: Songkhla Beach, 21. X. 2002, A. Freidberg, 4 ♂, 5 ♀ (all USNM); Saraburi; Ban Muak Lek Nat. Park, 6. VI. 1965, P. D. Ashlock, 9 ex., 5. VI. 1965, 1 ♂, 2 ♀ (BBM). TONGA: Niue I., 4–9. VII. 1963, M.R. Wheeler, 1 ♂; Tongatapu, Nukualofa, III. 1974, N.L.H. Krauss, 3 ♂, 3 ♀ and 25. X. 1945, D. G. Hall, 1 ♂ (all USNM), and 1 ♂ (BMNH); Vavau Neiafu, 0–100 m, III. 1974, N.L.H. Krauss, 13 ♂, 9 ♀ (USNM). UNITED ARAB EMIRATES: Sharjah Desert Park, light trap, 29. III.–6. IV. 2005, A. van Harten, 2 ♂, 3 ♀ (NMWC); Sweihan NARC, 14. III.–2.IV. 2005, light trap, A. van Harten, 1 ♀ (NMWC); Wadi Hayl, 6. III. 2010, T. Zatwarnicki, 1 ♀ (TZPC). VANUATU: Efate Vila, 0–100 m, II. 1974, N.L.H. Krauss, 1 ♂, 5 ♀ (CNC, USNM) and III. 1981, N. L. H. Krauss, 4 ♂, 1 ♀; Erromango, Dillon's Bay, II. 1981, N.L.H. Krauss, 4 ♂, 2 ♀; Tanna, Lenakel, III. 1981, N.L.H. Krauss, 4 ♂, 2 ♀ (all BMNH); Ile Vaté, Efate, 0–100 m, N. L. H. Krauss, 3 ♂, 2 ♀; Tagabe Ag. Sta., Port Vila, Efate, 3. VI. 1977, D. J. Bickel, 1 ♂ (CNC). VIETNAM: Bac Kan: Ba Be NP, Bo Lu, wet pasture, weeping, 22,3975976ºN, 105,6148598ºE, 202 m, 17. IV. 2010, L. Papp et al., 1 ♂; Ba Be NP, Na Mam, forest in the gorge behind the NP HQ, sweeping, 22,4438762ºN, 105,6638877ºE, 169 m, 17–18. IV. 2010, L. Papp et al., 1 ♂, 1 ♀; Na Kieng, along a muddy stream, sweeping, 22,4231128ºN, 105,6473557ºE, 222 m, 19. IV. 2010, L. Papp et al., 3 ♂, 2 ♀; Na Lang, along a rocky and muddy streamlet between rice paddies, sweeping, 22,4438762ºN, 105,6638877ºE, 169 m, 19. IV. 2010, L. Papp et al., 1 ♂, 1 ♀ (HNHM). Dac Lac: 19 km NE of Ban Me Thuot, 18. V. 1960, L. W. Quate, 1 ♀ (BBM); Hoa Binh: 3 km NW of Mai Chau, along a canal in paddy fields, sweeping, 20,691263ºN, 105,075127ºE, 229 m, 22. IV. 2010, L. Papp et al., 1 ♂; 6 km W of Tan Son, forest edge in a gorge, at light, 20,7417977ºN, 104,9401984ºE, 938 m, 22. IV. 2010, L. Papp et al., 1 ♂; Lao Cai: Hoang Lien NP, Tram Ton, forest edge and mud, sweeping, 22,3493723ºN, 103,7704565ºE, 1915 m, 8. IV. 2010, L. Papp, Z. Soltész, 1 ♀; 16–17. XII. 2008, L. Papp, 1 ♂; Hoang Lien NP, Thac Bac, rocky stream bed and mud, sweeping, 22,3358285ºN, 103,781337ºE, 1731 m, 12. IV. 2010, L. Papp et al., 1 ♂; Sin Chai, Hoang Lien NP, partly open slopes grazed by buffaloes, 22º20.434'N, 103º 49.349E, 1586 m, 14. XII. 2008, L. Papp, 1 ♂, 1 ♀ (HNHM).

Distribution ( Fig. 10 ): American Samoa, Australia (New South Wales, Northern Territory, Qeensland, Western Australia), Brunei, Burma (= Myanmar), Cambodia, China (Fukien), Fiji, Guam, India (Andhra Pradesh, Assam, Bengal, Karnataka, Kerala, Maharashtra, Tamil Nadu, Uttar Pradesh), Indonesia (Java), Laos, Malaysia ( Sarawak, Sabah), Micronesia (Yap), Nepal, New Caledonia, Northern Mariana Islands, Oman, Palau Islands, Papua New Guinea (including Bismarck Isl.), Philippines, Samoa, Solomon Islands, Sri Lanka, Taiwan, Thailand, Tonga, United Arab Emirates, Vanuatu, Vietnam.

Remarks. Externally this species is very similar to L. africana but is distinguished from this species by the black basiflagellomere and palpus.

Figures

FIGURE 7. Head of Leptopsilopa mutabilis Cresson. A. Anterior view. B. Oblique view.

FIGURE 8. Structures of the male terminalia of Leptopsilopa pollinosa (Kertész) (Sri Lanka: Maskellya Oya). A, Epandrium, cerci, and pregonostyli, ventral view. B, Same, lateral view. C, Internal genitalic structures (aedeagus (shaded), phallapodeme, medandrium, postgonostylus, gonite, and hypandrium), ventral view. D, Same, lateral view. Scale bar 0.1 mm.

FIGURE 9. Head of Leptopsilopa pollinosa (Kertész). A. Anterior view. B. Oblique view.

FIGURE 10. Distribution map for Leptopsilopa mutabilis Cresson (circles) and L. pollinosa (Kertész) (hatched area).