Anelaphus pilosus Chemsak & Noguera, 2003

Santos-Silva, Antonio, 2021, Description of four new species of Anelaphus Linsley (Coleoptera, Cerambycidae Cerambycinae, Elaphidiini), correction of the spelling of species-group names and description of the female of Anelaphus pilosus Chemsak and Noguera, Zootaxa 5048 (3), pp. 371-390 : 385-387

Anelaphus pilosus Chemsak & Noguera, 2003

( Figs 40–45 )

Anelaphus pilosus Chemsak & Noguera, 2003: 62 ; Monné, 2005: 196 (cat.); Maes et al., 2010: 168 (distr.); Monné, 2021: 309 (cat.).

Description. Female ( Figs 40–43 ). Ventral surface of head dark brown; dorsal surface of head black; parts of ventral mouthparts dark reddish brown; apex of maxillary and labial palpomeres yellowish brown; anteclypeus brown; scape, pedicel and antennomeres III–V dark reddish brown with some areas darkened; antennomeres VI–XI reddish brown. Pronotum and sides of prothorax mostly black. Ventral surface of thorax mostly dark brown with some areas black, especially on metathorax. Elytral suture and epipleural margin black; remaining surface dark reddish brown (more dark brown depending on light intensity. Legs dark reddish brown with some areas darkened. Ventral surface of abdomen black basally, gradually dark brown toward apex.

Head. Frons coarsely, abundantly punctate, except nearly smooth central place close to postclypeus; on wide central area with somewhat abundant, decumbent yellowish-white setae, not obscuring integument, and long erect yellowish-brown setae interspersed; area close to eyes with dense yellowish-white setae, part of them yellowishbrown apically, and long, erect yellowish-brown setae interspersed. Area between antennal tubercles and upper eye lobes coarsely, abundantly punctate, except smooth central area between eyes; remaining surface of vertex densely, shallowly, confluently punctate; area between antennal tubercles with setae as on frons; area between antennal tubercles and anterior margin of upper eye lobes with somewhat abundant, both yellowish-white and yellowishbrown setae, almost forming a transverse band, distinctly denser laterally, gradually sparser toward central area; area between upper eye lobes with abundant yellowish-white setae laterally, not obscuring integument, gradually sparser toward central area; remaining surface of vertex mostly glabrous; vertex tumid from posterior margin of eyes to prothorax. Area behind upper eye lobes coarsely, shallowly, confluently punctate; with somewhat sparse yellowishwhite setae close to eyes, glabrous on remaining surface. Area behind lower eye lobes smooth, glabrous close to eye (this area gradually widened toward ventral surface), except short yellowish-white setae close to inferior area of eye; remaining surface; area close to prothorax coarsely, confluently punctate, with a few short yellowish-white setae. Genae with yellowish-white pubescence not obscuring integument, and yellowish-brown setae interspersed, except glabrous distal area. Wide central area of postclypeus transversely carinate close to frons; with bristly, short yellowish-brown setae centrally, and long, erect setae of same color laterally. Sides of postclypeus smooth, glabrous. Labrum subcoplanar with anteclypeus at posterior half, inclined at anterior half; with minute, yellowish-white setae on subcoplanar area, and long, erect yellowish-brown setae on inclined area (setae longer laterally). Gulamentum smooth, glabrous on posterior half; anterior half coarsely rugose-punctate; with short, erect yellowish setae (appearing to be whiter depending on light intensity), and long, erect setae of same color interspersed. Posterior 2/3 of outer side of mandibles coarsely rugose-punctate, with short, sparse yellowish-white setae, and long, erect, thick yellowishbrown setae interspersed; anterior third smooth, glabrous. Lower eye lobes with short, erect yellowish-white setae among the ommatidia. Distance between upper eye lobes 0.25 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.40 times distance between outer margins of eyes. Antennae 1.4 times elytral length, almost reaching elytral apex. Scape finely, densely punctate, except smooth dorsal apex; with abundant yellowish-white pubescence partially obscuring integument, except glabrous smooth area, and long, erect setae yellowish setae interspersed. Pedicel and antennomeres III–XI with abundant yellowish-white pubescence not obscuring integument, pubescence shorter and denser from VII; antennomeres III–VI with long, erect yellowishbrown setae ventrally (gradually sparser toward VI); antennomeres III–X with long, erect yellowish-brown setae apically; antennomeres III–VI with short spine on inner apex (spine gradually shorter toward VI); outer apical angle of antennomeres III–IV rounded; outer apical angle of antennomeres V–X angular; apical third of antennomere V and entire outer side of antennomeres VI–X gradually flattened from about middle to outer margin (flattened area densely, microscopically punctate). Antennal formula based on length of antennomere III (excluding apical spines): scape = 1.37; pedicel = 0.25; IV = 0.87; V = 1.10; VI = 1.03; VII = 1.00; VIII = 0.97; IX = 0.88; X = 0.80; XI = 0.90.

Thorax. Prothorax wider than long; sides divergent from anterolateral angles to middle, convergent from middle to posterolateral angles. Pronotum coarsely, densely, punctate (punctures confluent on some areas) except smooth central area on posterior third; with abundant yellowish-white pubescence not obscuring integument, denser on sides of anterior half, absent on smooth area (pubescence appearing to be whiter or grayish-white depending on light intensity and angle source); with a few long, erect yellowish setae interspersed. Sides of prothorax coarsely, abundantly punctate, except nearly smooth anterior and posterior areas; with abundant yellowish-white pubescence not obscuring integument, and long, erect yellowish setae interspersed, except almost glabrous anterior and posterior areas. Prosternum coarsely, mostly shallowly, partially confluently punctate on posterior 3/4, finely, shallowly, sparsely punctate on anterior quarter; with abundant yellowish-white pubescence on posterior 3/4 (appearing to be whiter depending on light intensity), almost glabrous on anterior quarter; with long, erect, sparse yellowish setae throughout. Prosternal process with dense yellowish-white pubescence obscuring integument, except apex and sides of posterior widened area with sparse yellowish-brown pubescence; narrowest area of prosternal process 0.08 times procoxal width. Procoxal cavities open behind (opening width smaller than apical width of prosternal process). Anterocentral region of mesoventrite depressed on each side of middle. Ventral surface of meso- and metathorax with abundant yellowish-white pubescence (appearing to be whiter depending on light intensity), except glabrous basal area of center of mesoventrite, sides of basal 3/4 of central area of mesoventrite with sparse pubescence, and glabrous metathoracic discrimen; metanepisternum and metaventrite with long, erect, sparse yellowish-brown setae. Scutellum with abundant yellowish-white pubescence, except glabrous anterocentral region. Elytra. Coarsely, abundantly punctate on anterior third, punctures gradually finer toward apex; apex narrowly subtruncate (appearing to be oblique depending on viewing angle), with sutural angle slightly projected; with abundant yellowish-white pubescence not obscuring integument (appearing to be whiter depending on light intensity), and long, erect, somewhat abundant yellowish-brown setae interspersed. Legs. Femora with abundant yellowish-white pubescence not obscuring integument (appearing to be whiter depending on light intensity); femora with long, yellowish-brown setae interspersed, more abundant dorsally on metafemora. Tibiae with abundant yellowish-white pubescence not obscuring integument (appearing to be whiter depending on light intensity), except bristly, dense, yellowish-brown pubescence on ventral surface of posterior half of protibiae and ventral surface of posterior third of meso- and metatibiae. Metatarsomere I shorter than II–III together.

Abdomen. Ventrites with abundant yellowish-white pubescence partially obscuring integument, and long yellowish-brown setae interspersed. Apex of ventrite 5 slightly rounded.

Dimensions (mm). Total length, 13.60; prothoracic length, 2.70; anterior prothoracic width, 2.10; posterior prothoracic width, 2.65; maximum prothoracic width, 3.10; humeral width, 3.85; elytral length, 9.30.

Material examined. GUATEMALA (new country record), Zacapa: Quarry Rd to San Lorenzo, 15.043871, -89.668775, Sierra Las Minas , 600 m, 1 female, 26.V.2019, Skillman, Wappes and Monzón leg. ( FWSC) .

Remarks. According to Chemsak & Noguera (2003) the antennae in the holotype male are 12-segmented, although the 12 th antennomere is missing. The female described here has antennae distinctly 11-segmented. However, in Aneflus LeConte, 1873 , when the antennae are 12-segmented only the male has this feature, while the female has antennae 11-segmented (see, for example, A. minutivestis Chemsak & Linsley, 1963 ). According to Chemsak (1959), the antennae in males of Anelaphus antennatus Chemsak, 1959 are 12-segmented. Hovore (1983) synonymized A. antennatus with A. subinermis Linsley, 1957 , and reported: “In 27 male subinermis collected near Fort Davis, the eleventh segment displays an allometric tendency for appendiculation at the apical one-third, ranging from a feeble transverse sulcus in smaller specimens to a complete division (a “twelfth” segment) in very large individuals.” If the synonymy proposed by Hovore (1983) is right (the holotype male of A. subinermis is a large specimen and shows no trace of division in the antennomere XI), then it is possible that the same variation also occurs in males of A. pilosus . Photographs of the holotype male of A. pilosus ( Figs 44–45 ) sent by Peter T. Oboyski (EMEC) show that the antennae are really 12-segmented and that the last segment is missing (the 11 th segment is not broken apically).

Figures

FIGURES 40–45. Anelaphus pilosus Chemsak and Linsley, 2003. 40–43) Female: 40) Dorsal habitus; 41) Ventral habitus; 42) Lateral habitus; 43) Head, frontal view. 44–45) Holotype male: 44) Lateral view, left side; 45) Lateral view, right side.