Genieridium bidens ( Balthasar, 1938 ) Vaz-De-Mello, 2008

Vaz-De-Mello, Fernando Z., 2008, Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus, Zootaxa 1955, pp. 1-75 : 28-29

6.1. Genieridium bidens ( Balthasar, 1938) , new combination

Pedaridium bidens: Balthasar 1938: 218–220 ; Blackwelder 1944: 203; Vaz-de-Mello & Génier 2005: 45–46

Pedaridium hirsutum: Pessôa & Lane 1941: 437

Pedaridium brasiliensis Ferreira & Galileo, 1993 : Ferreira & Galileo 1993: 5–7, 21–23, 54; Koller et al. 1999: 405, 407; Vaz-de-Mello 2000: 194; Verdú & Galante 2001: 599–601

Type series: Pedaridium bidens Balthasar, 1938 . Lectotype (designated by Vaz-de-Mello & Génier, 2005): ♂, Paraguay ( NMP). Paralectotypes: BRAZIL: Goiás: Jatahy (1 NMP, 1 NHMB)

Pedaridium brasiliensis Ferreira & Galileo, 1993 : Holotype: BRAZIL: Bahia: Encruzilhada , 980 m, XI- 1972, Alvarenga (MZSP)

Paratypes: BRAZIL: Bahia: Encruzilhada , 980 m, XI-1972, Alvarenga (8 MZSP) ; Mato Grosso do Sul: Murtinho (luz), I-1930, R Spitz (2 MZSP) ; Três Lagoas , Faz. Dr. José Mendes, 15-30-V-1964, Exp. Dep. Zool. (1 MZSP) ; Faz. Retiro das Telhas, 15-30-V-1964, Exp. Dep. Zool. (2 CMNC, 11 MZSP) ; Mato Grosso: Chapada dos Guimarães , XI-1963, Alvarenga (3 MZSP) ; Minas Gerais: Arinos , 06-08-XI-1964, Exp. Dep. Zool. (2 MZSP) ; Paraná: Vila Velha , XI-1944, Hatsbach (1 CMNC, 1 MZSP) ; São Paulo : Castilho, marg. esq. Rio Paraná , X-1964, Exp. Dep. Zool. (1 MZSP) ; Franca , VIII-1910, Garbe (2 MZSP) ; Itu, Faz Pau d'Alho , 28-29-X-1965, Martins & Biasi (1 CMNC) ; Itú , 27-XII-1957, U. Martins (4 MZSP) ; XI-1958, U. Martins (2 MZSP) ; Pirassununga , 09-X-1945, Schubart (1 MZSP) .

Non-type material examined: BRAZIL: Bahia: no data, Bondar (1 MNRJ) ; Encruzilhada , 980 m, XI- 1972, Alvarenga (1 IBSP) ; XI-1974, Alvarenga (1 CMNC) ; XII-1980, Alvarenga & Martínez (8 CMNC) ; Vitória da Conquista , I-1993 (2 MZSP) ; Distrito Federal: Est. Florestal Cabeça do Veado , 1100 m, X-1971, EG, I & EA Munroe (1 CNCI) ; Goiás: Bom Jesus , X-1996, J Carlos (1 FVMC) ; Niquelândia , X-1993, arm luminosa (1 FVMC) ; Rio Verde , 17-XI-1984, Bello (1 FVMC) ; X-1992, J Carlos (2 AMBC) ; XI-1993 (1 AMBC) ; XI-1995 (2 AMBC) ; Mato Grosso do Sul: Costa Rica , 17-XII-1993, S Ide (2 MZSP) ; Murtinho , XII-1929, Malkin (1 MNRJ) ; Selvíria , UNESP farm, 01-V-1990, CAH Flechtmann, ex Guzerá bovine dropping, Brachiaria decumbens pasture (1 FEIS) ; 01-VI-1991 (2 FEIS) ; 06-VII-1991 (1 FEIS) ; 06-VIII-1991 (1 FEIS) ; 13-XII-1992 (1 FEIS) ; 15-VI-1991 (2 FEIS) ; 16-II-1991 (1 FEIS) ; 18-III-1999, ex black light (2 FEIS) ; 18-V-1991, ex Guzerá bovine dropping (1 FEIS) ; 20-I-1999, ex black light (24 FEIS) ; 20-VIII-1991, ex Guzerá bovine dropping (1 FEIS) ; 20-X-1991 (3 FEIS) ; 22-IV-1992 (2 FEIS) ; 23-II-1991 (3 FEIS) ; 23-II- 1992 (1 FEIS) ; 27-IV-1991 (1 FEIS) ; 31-X-1993 (1 FEIS) ; Terenos , 01-IV-1994, WW Koller (1 FVMC) ; Mato Grosso: Chapada dos Guimarães , XI-1963, Alvarenga (1 MZSP) ; Minas Gerais: Águas Vermelhas , XII-1997, Bello (3 AMBC) ; XII-1998, Bello & Vaz-de-Mello (1 FVMC) ; Cordisburgo, Faz Pontinha, X- 1993, FZ Vaz-de-Mello (1 CMNC, 2 BDGC); Ipatinga , XI-1992, E Grossi (1 FVMC) ; Montes Claros , I-2000, JNC Louzada (1 FVMC) ; Três Marias , X-1989 (2 FVMC) ; Pará: Belém , IX-1964, E Dente (1 MZSP) ; Canindé ( Rio Gurupi ), X-1964, Malkin (1 CMNC) ; São Paulo : Bálsamo , 12-XI-1987, C Bergmann, seringueira (1 IBSP) ; 12-XII-1987 (1 IBSP) ; 13-X-1988 (1 IBSP) ; 19-II-1987 (1 IBSP) ; 29-X-1987 (1 IBSP) ; Castilho , marg. esq. Rio Paraná, 15-22-IX-1962, Exp. Dep. Zool. (1 IBSP) ; Itirapina , 15-IX-1996, JR Verdú (2 FVMC) ; Itu, Faz Pau d'Alho , I-1959, Martins (1 CMNC) ; Mirante do Paranapanema , 09-X-1991, J Rodrigues, black light, pasture area (2 FEIS) ; 13-XI-1991 (2 FEIS) ; Pradópois , XII-1976, PM Botelho (1 FVMC, 4 ESAP); São Carlos , 02-XII-1993, J Mendes (1 FVMC) ; 11-XI-1993 (1 FVMC) ; PARAGUAY: no data: (1 HMNH) ; Anisits (1 HMNH) ; Boquerón: Pto Casado , XI-1950, Martínez (2 CMNC) ; Caaguazú: Caaguazú, Piscicultura , II-1986, A Martínez (1 BDGC, 6 CMNC); Central : Asunción, 10-X-1904, Vezényi (1 HMNH) ; 28-IX- 1904 (1 HMNH) ; Concepción: Horquetá , XII-1950, Martínez, a la luz (6 CMNC) ; Paraguary: Sapucay , 05- XI-1991, U Drechsel (9 ABC) ; San Pedro: Cororó , XI-1999, G Arriágada (1 FVMC) ; Rio Ypane, Cororó , XI-1979, A Martínez (1 BDGC, 19 CMNC) .

Diagnosis: 3.8–5.3 mm. Surface dull. Clypeal teeth obtuse and equilateral, continuous (at same plane) with clypeal margin, separated by wide V-shaped emargination. Eyes, in dorsal view, visible and very small (Fig. 55). Elytral striae lacking punctures. Very similar to G. margareteae , but lacking punctures in striae, and anterior clypeal carina. Parameres as in Fig. 56.

Distribution: The entire Cerrado province of the Chacoan subregion, with invasion of pasturelands in neighboring provinces ( Fig. 104).

Remarks: This is a common pastureland species in Brazil, and may be related to G. margareteae .