Antillocladius Saether

Humberto F. Mendes, Trond Andersen & Ole A. Saether, 2004, A review of Antillocladius Saether, 1981; Compterosmittia Saether, 1981 and Litocladius new genus (Chironomidae, Orthocladiinae), Zootaxa 594, pp. 1-82 : 25-26

Key to the males of Antillocladius Saether

1. Squama bare ( Fig. 16 D); Mexico..................................... A. zempoalensis new species

— Squama with setae ......................................................................................................... 2

2. Anal point nearly parallel­sided with numerous, weak, partly anterior directed setae ; inferior volsella either pointed triangular or consisting of a long apically simple or bifid lobe; wing bare ..................................................................................................... 3

— Anal point tapering with numerous, usually strong and posterolateral directed setae ; inferior volsella either with a dorsal anterior triangular to digitiform part and a more rounded ventral lobe or circular with or without additional rounded posterior extension and conspicuously set off; wing usually with apical setae ........................................... 6

3. Inferior volsella pointed triangular, costal extension very long................................... 4

— Inferior volsella a long posteromedially directed simple or apically bifid lobe, costal

extension of moderate length ....................................................................................... 5 4. Inferior volsellae triangular, leaf­like ( Fig. 5 D); Brazil............... A. folius new species

— Inferior volsella small, sharply pointed ( Fig. 3 D); Brazil............. A. biota new species

5. Inferior volsella apically bifid ( Fig. 13 C); Brazil................ A. sooretama new species

— Inferior volsella simple ( Fig. 14 D); Brazil, Venezuela............ A. ubatuba new species

6. Virgaatleastnearlyaslongashalfphallapodemelength; wingmembranewithapicalsetae ....................................................................................................................................... 7

— Virga absent or short; wing membrane with or without apical setae .......................... 11

7. Anal point with numerous weak setae ; inferior volsella weak ( Fig. 15 D); Venezuela, Ecuador....................................................................... A. venequatoriensis new species

— Anal point with strong, stiff setae ; inferior volsella well developed............................ 8

8. Wing with few setae apically in cell r4+5; AR 1.4–1.7................................................. 9

— Wing with more numerous apical setae present also in cells m1+2 and sometimes m3+4; AR 1.0–1.6 ................................................................................................................. 10

9. Virga nearly as long as phallapodeme ( Fig. 9 E); Brazil.............. A. musci new species

— Virga less than half as long as phallapodeme; China, Thailand...................................... ........................................................................................... A. zhengi Wang and Saether

10. Virga about 2/3 as long as phallapodeme; squama with 5–6 setae ; cell m3+4 with sev­ eral setae ; Ecuador, Mexico, Nicaragua, USA............................ A. pluspilalus Saether

— Virgahalfaslongasphallapodeme;squamawith 8–13 setae ;cellm3+4 bare; China, Russia................................................................................... A. scalpellatus Wang and Saether

11. Virga consisting of 2 or 6 very short spines; costal extension short .......................... 12

— Virga absent; costal extension long............................................................................ 13

12. Virga consisting of 6 spines; inferior volsella with digitiform dorsal anterior lobe cov­ ering rounded posterior, ventral lobe ( Fig. 4 F); Mexico A. calakmulensis new species

— Virga consisting of 2 spines; inferior volsella apically divided into 2 semicircular lobes ( Fig. 8 F); Mexico................................................................ A. herradurus new species

13. Inferior volsella a well set off circular lobe with additional posterior semicircular extension; wing membrane bare; Brazil, Mexico, Venezuela, USA A. arcuatus Saether

— Inferior volsella a well set off circular lobe; wing membrane usually with setae ...... 14

14. Wing membrane with numerous apical setae , present also in cell m3+4; Ecuador.......... ................................................................ A. skartveiti Andersen and Contreras­Ramos

— Wing membrane with 0–3 setae apically in cell r4+5; Brazil, Saint Vincent, Venezuela...................................................................................................... A. antecalvus Saether

Figures

FIGURE 3. Antillocladius biota new species, male imago. A – cibarial pump, tentorium and stipes; B – thorax; C – wing; D – anal point and tergite IX and dorsal aspect of left gonocoxite; E – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect; F – gonostylus.

FIGURE 4. Antillocladius calakmulensis new species, male imago. A – head; B – cibarial pump, tentorium and stipes; C – thorax; D – wing; E – anal point and tergite IX and dorsal aspect of left gonocoxite; F – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect; G – gonostylus.

FIGURE 5. Antillocladius folius new species, male imago. A – cibarial pump, tentorium and stipes; B – thorax; C – wing; D – anal point and tergite IX and dorsal aspect of left gonocoxite; E – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect; F – gonostylus.

FIGURE 8. Antillocladius herradurus new species, male imago. A – head; B – cibarial pump, tentorium and stipes; C – thorax; D – wing; E – anal point and tergite IX and dorsal aspect of left gonocoxite; F – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect; G – gonostylus.

FIGURE 9. Antillocladius musci new species, male imago. A – cibarial pump, tentorium and stipes; B – thorax; C – wing; D – anal point and tergite IX and dorsal aspect of left gonocoxite; E – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect; F – gonostylus.

FIGURE 13. Antillocladius sooretama new species, male imago. A – cibarial pump, tentorium and stipes; B – wing; C – anal point and tergite IX and dorsal aspect of left gonocoxite; D – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect; E – gonostylus.

FIGURE 14. Antillocladius ubatuba new species, male imago. A – cibarial pump, tentorium and stipes; B – thorax; C – wing; D – anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus; E – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect.

FIGURE 15. Antillocladius venequatoriensis new species, male imago. A – cibarial pump, tentorium and stipes; B – thorax; C – wing; D – anal point and tergite IX and dorsal aspect of left gonocoxite; E – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect; F – gonostylus.

FIGURE 16. Antillocladius zempoalensis new species, male imago. A – head; B – cibarial pump, tentorium and stipes; C – thorax; D – wing; E – anal point and tergite IX and dorsal aspect of left gonocoxite; F – hypopygium with anal point and tergite IX removed, left dorsal aspect, right ventral aspect.