Helochares songi, Jia & Tang, 2018
publication ID |
https://doi.org/ 10.5852/ejt.2018.438 |
publication LSID |
lsid:zoobank.org:pub:00B57E96-225C-4646-9CB8-B3514C4EC7E5 |
DOI |
https://doi.org/10.5281/zenodo.3794615 |
persistent identifier |
https://treatment.plazi.org/id/2B1792C1-5D59-46B2-AE98-3440BA2FED57 |
taxon LSID |
lsid:zoobank.org:act:2B1792C1-5D59-46B2-AE98-3440BA2FED57 |
treatment provided by |
Valdenar |
scientific name |
Helochares songi |
status |
sp. nov. |
Helochares songi View in CoL sp. nov.
urn:lsid:zoobank.org:act:2B1792C1-5D59-46B2-AE98-3440BA2FED57
Figs 1–8 View Figs 1–8 , 48–49 View Figs 47–55
Diagnosis
Length 4.8–5.3 mm, width 2.6–2.9 mm. Dark brown or black ( Fig. 1 View Figs 1–8 ), clypeus uniformly dark brown to black ( Fig. 4 View Figs 1–8 ), anterior half of lateral margin of pronotum yellowish brown ( Figs 1, 3–4, 6 View Figs 1–8 ), elytra uniformly dark brown to black ( Fig. 3 View Figs 1–8 ). Dorsum with uniform dense and coarse ground punctures. Clypeus moderately expanded laterally, with somewhat rounded lateral margin, clearly impressed in front of eyes ( Fig. 4 View Figs 1–8 ). Maxillary palps ca 1.2× as long as width of head, dark brown to black, each palpomere with light colour apically ( Figs 1–2, 4 View Figs 1–8 ). Pedicel subequal in length to antennomeres 3–6 combined. Submentum with a small glabrous area medially and with a few coarse punctures laterally ( Fig. 5 View Figs 1–8 ). Elytra almost parallel from base to posterior third, somewhat explanate laterally, with distinct systematic punctures ( Fig. 3 View Figs 1–8 ). Mesoventrite with a low transverse ridge medially and a low longitudinal carina ( Fig. 7 View Figs 1–8 ). Metaventrite without glabrous area medially. Femora densely pubescent except for apical eighth ( Figs 2, 8 View Figs 1–8 ). Aedeagus slender, length of paramere ca 10 × as long as width of apex ( Fig. 48 View Figs 47–55 ), subparallel, apex with a small tooth inwards ( Figs 48–49 View Figs 47–55 ); apex of median lobe slightly overlapping parameres; basal strut of median lobe long, ca ¼ × as long as parameres ( Fig. 48 View Figs 47–55 ).
Etymology
The specific name is given in honor of Keqing Song, the collector of the type specimens.
Material examined
Holotype
CHINA: 1 ♂, Guangxi Province, Shiwandashan, Nalin River , 316 m a.s.l., in water at edge of upper river, 21 Jul. 2011, Keqing Song leg. ( SYSU).
Paratypes (77 spec., BMNH, KMNH, NMPC, SYSU)
CHINA: Guangxi: 2 ƋƋ, 4 ♀♀, 15 spec., same collection data as for holotype; 3 spec., Shiwandashan, Forest Park, 239 m a.s.l., edge of upper river, 19 Jul. 2011, Keqing Song leg.; 11 spec., Shiwandashan, Forest Park, edge of river, 17 Aug. 2011, Keqing Song leg.; 3 spec., Shiwandashan, Forest Park, 221 m a.s.l., in water at edge of lower stone river, 12 Jul. 2011, Keqing Song leg.; 20 spec., Shiwandashan, 430 m a.s.l., edge of Nalinhe River (in water), 21 Jul. 2011, Keqing Song leg.; 19 spec., Shiwandashan, 269 m a.s.l., edge of river outside Forest Park, 20 Jul. 2011, Keqing Song leg. ( BMNH, KMNH, NMPC, SYSU).
Description
FORM AND COLOUR. Body oval, length 4.8–5.3 mm, width 2.6–2.9 mm, moderately convex. Head dark brown to black. Pronotum dark brown to black, anterior half with narrow yellowish brown lateral margins ( Figs 1, 3–4, 6 View Figs 1–8 ). Elytra uniformly dark brown to black ( Figs 1, 3 View Figs 1–8 ). Maxillary palps dark brown to black, each palpomere light yellowish brown apically ( Figs 1–2, 4 View Figs 1–8 ). Labial palpomeres yellow to yellowish brown. Antennae yellowish brown, club with dense yellow setae. Ventral side dark brown or black ( Fig. 2 View Figs 1–8 ), legs with same colour as ventral side, but with reddish to yellowish brown tarsomeres ( Fig. 2 View Figs 1–8 ).
HEAD. Labrum with fine and dense punctures, smooth between punctures, anterior margin broadly emarginate. Clypeus broad, anterior margin clearly broadly emarginate, moderately expanded in front of eyes, with somewhat rounded lateral margin ( Fig. 4 View Figs 1–8 ), clearly impressed in front of eyes ( Fig. 4 View Figs 1–8 ). Ground punctures on clypeus coarser and sparser than those on labrum, surface smooth between punctures. Systematic punctures on clypeus almost same size as ground punctures. Eyes of moderate size, clearly emarginate anteriorly ( Figs 1, 4 View Figs 1–8 ), separated by ca 4.0–4.1 × the width of one eye. Frons with slightly coarser and stronger punctures than on clypeus, with clear systematic punctures. Maxillary palps long and slender, ca 1.1–1.2 × as wide as head, second and third palpomeres curved inwards, apical palpomere almost as long as penultimate, asymmetrical ( Fig. 4 View Figs 1–8 ). Antennae with 9 antennomeres, scape ca 1.5× as long as pedicel, pedicel subequal in length to antennomeres 3–6 combined, club loosely segmented, with dense pubescence. Mentum ca as wide as long, subquadrate, with deep v-shaped impression anteromedially, with coarse punctures and wrinkles posteriorly ( Fig. 5 View Figs 1–8 ). Submentum with a small glabrous area medially and with a few coarse punctures laterally. Maxilla with coarse punctures, smooth between punctures.
THORAX. Pronotum ca 2× as wide as long, widest posteriorly, posterolateral margins broadly round, anterior margin smooth, with a very fine transverse groove, lateral margin with stronger groove, posterior margin without such groove; posterior margin almost straight. Ground punctures slightly coarser than those on head, systematic punctures on pronotum distinct, almost of same size as ground punctures ( Figs 4, 6 View Figs 1–8 ). Prosternum bulging in middle, not carinate, pubescent, with transverse groove ( Fig. 5 View Figs 1–8 ), protruding anteriorly. Mesoventrite with a low transverse ridge medially, which does not bear an elevated tooth or projection but with long setae ( Fig. 7 View Figs 1–8 ); behind ridge with a low longitudinal carina. Metaventrite pubescent, with somewhat convex middle portion, without glabrous area. Metepisterna ca 3.5× as long as wide, subparallel. Scutellum triangular, with similar punctures to elytra. Elytra almost parallel from base to posterior third, somewhat explanate laterally, ground punctures similar to those on pronotum, with 3 distinct rows of systematic punctures ( Fig. 3 View Figs 1–8 ), without sutural stria; epipleuron very broad anteriorly, reaching posterior margin of first visible abdominal sternite, with sparse, coarse punctures. Femora densely pubescent, with apical eighth glabrous ( Figs 2, 8 View Figs 1–8 ). Metatarsomeres with dense white hairs ventrally and a few long swimming hairs dorsally. Claws of moderate size, rather strongly curved.
ABDOMEN. All visible abdominal ventrites with dense pubescence; first ventrite not carinate ( Figs 2, 8 View Figs 1–8 ).
AEDEAGUS. Slender, length of parameres ca 10 × as long as width of apex, subparallel, a little expanded subapically and round apically ( Fig. 48 View Figs 47–55 ), apex with a small, inwardly directed tooth ( Figs 48–49 View Figs 47–55 ); median lobe slightly protruding beyond parameres, gradually narrowed apically, apex ca 0.5× as wide as apex of parameres; basal strut long, ca ¼ × as long as parameres ( Fig. 48 View Figs 47–55 ).
Differential diagnosis
This species is very similar to H. fuliginosus d’Orchymont, 1932 , but may easily be distinguished from the latter by its larger size (3.6–4.3 mm in H. fuliginosus ), lateral margin of clypeus and elytra with same colour as disc ( Fig. 4 View Figs 1–8 ), pronotum and elytra more coarsely punctate, and maxillary palps dark brown or black, each palpomere with light colour apically ( Figs 4–5 View Figs 1–8 ). Aedeagus slender, parameres ca 10 × as long as width of apex ( Fig. 48 View Figs 47–55 ), apex with a small inwardly directed tooth ( Figs 48–49 View Figs 47–55 ); median lobe slightly projecting beyond parameres apically ( Fig. 48 View Figs 47–55 ). It can be distinguished from H. atropiceus Régimbart, 1903 , H. ciniensis Hebauer, 1999 and H. taprobanicus Sharp, 1890 , occurring in the Oriental, by its slender aedeagus, parameres ca 10× as long as width of apex, without branches, not concave outside, apex with a small inwardly directed tooth ( Figs 48–49 View Figs 47–55 ). Hebauer (1995) described Helochares fulgurans based on a unique female from Thailand. Helochares songi sp. nov. can be distinguished from H. fulgurans by its larger size (3.5 mm in H. fulgurans ), dark colour (bright yellow brown, only dark between eyes, with two dark basal spots on pronotum in H. fulgurans ) and maxillary palps dark brown or black, each palpomere with light colour apically (yellow in H. fulgurans ). From H. vitalisi d’Orchymont, 1919 ( Fig. 45 View Figs 41–46. – 41–42 ) it differs in having the head, pronotum and elytra with coarser punctures (ground punctures very fine in H. vitalisi ), each maxillary palpomere with light colour apically, clypeus moderately expanded laterally, with somewhat rounded lateral margin, clearly impressed in front of eyes (clypeus not expanded laterally and not impressed in front of eyes in H. vitalisi ) and elytra almost parallel from base to posterior third (widest at middle in H. vitalisi ), somewhat explanate laterally (not explanate in H. vitalisi , as in Enochrus ).
Biology
Lives along the sides of rivers with a sandy bottom and stagnant water. The female carries the egg case under the abdominal ventrites.
Remarks
The density is very high at the type locality, but no specimen was collected at light in July. It is probable that this species can be collected at light during its reproductive period.
Distribution
China (Guangxi), so far only known from the Shiwandashan National Park and its surroundings.
NMPC |
National Museum Prague |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Acidocerinae |
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