Simplomys julii, (DAAMS, 1989)

García-Paredes, Israel, Peláez-Campomanes, Pablo & Álvarez-Sierra, M. Ángeles, 2009, Gliridae (Rodentia, Mammalia) with a simple dental pattern: a new genus and new species from the European Early and Middle Miocene, Zoological Journal of the Linnean Society 157 (3), pp. 622-652 : 641-642

SIMPLOMYS JULII ( DAAMS, 1989) ( FIG. 7 )

Synonymy: Pro-parte Peridyromys murinus from Villafeliche 2A in Daams (1981).

Original diagnosis ( Daams, 1989: 16): ‘A small Pseudodryomys species with a flat to slightly convex occlusal surface. The M1,2 have a subquadrate circumference. Simple dental pattern. Centrolophs are absent, and centrolophids are short or absent. Protoloph and metaloph form a Y – pattern in M1,2’.

Emended diagnosis: Small Simplomys species with molars that have a flat to slightly convex occlusal surface and simple pattern. Anteroloph and posteroloph are situated slightly below the plane of the trigon (protoloph–metaloph). The protoloph and metaloph form a composite ‘Y’ shape with a bent connection. A line connecting the labial end of the metaloph and the lingual end of the protoloph is aligned at right angles to the anteroposterior axis of the tooth. The occlusal surface of the M1 and M2 has a square outline. In M1 the anteroloph and the anterior valley are straight and parallel to the anterior border of the tooth. In the lower molars, the centrolophid is absent or poorly developed and it does not extend into the central valley. The m3 has the posterior part reduced in size and in most cases the posterior valley is absent.

Differential diagnosis: Simplomys julii differs from S. simplicidens , S. robustus , and S. aljaphi in its smaller size, less concave cheek teeth, and by having the centrolophid less developed.

Type locality: Artesilla , Calatayud–Montalbán basin, Spain (local zone C, MN 4 , Lower Miocene) .

Stratigraphical distribution: Local zones A to Db (MN 3-MN 4, Lower and Middle Miocene).

Geographical distribution: Spain and France (see Supporting Information Appendix S1).

Figures

Figure 7. Simplomys julii (Daams, 1989) from Artesilla (Fig. 7A–H; J–L; N–O) and San Roque 3 (Fig. 7I, M) (collection MNCN). A, P4 ART-2444. B, M1 ART-3. C, M2 ART-28. D, M3 ART-21. E, P4 ART-2447. F, M1 ART-4 holotype. G, M2 ART-13. H, M3 ART-1176. I, p4 SR3-803. J, m1 ART-16. K, m2 ART-14. L, m3 ART-1036. M, p4 SR3-604. N, m1 ART-22. O, m2 ART-27. Right side specimens underlined. Holotype in italics.