Simplomys robustus, (DE BRUIJN, 1967)

García-Paredes, Israel, Peláez-Campomanes, Pablo & Álvarez-Sierra, M. Ángeles, 2009, Gliridae (Rodentia, Mammalia) with a simple dental pattern: a new genus and new species from the European Early and Middle Miocene, Zoological Journal of the Linnean Society 157 (3), pp. 622-652 : 640

SIMPLOMYS ROBUSTUS ( DE BRUIJN, 1967) ( FIG. 5 )

Original diagnosis ( De Bruijn, 1967: 259, translated from Spanish): ‘[Large Pseudodryomys with simple dental pattern. The upper molars have neither centrolophs nor extra ridges. The m1 and m2 have a small centrolophid. The m3 is reduced]’.

Emended diagnosis by Daams (1974: 236): ‘ Pseudodryomys -assemblages consisting of large teeth which differ from P. simplicidens in size and in the less reduced m3. The dental patterns of the other molars are identical of those of P. simplicidens ’.

Emended diagnosis: Large Simplomys in which the dental pattern in m1 and m2 is formed by the four main ridges and a well-developed centrolophid. The latter extends diagonally into the central valley but never exceeds the middle part of the occlusal surface. The m3 always has a posterior valley and shows little reduction in its posterior part. The M1 may rarely present a small anterior centroloph.

Differential diagnosis: Simplomys robustus differs from S. simplicidens by having the premolars and the m3 less reduced in size and less simplified in morphology. The posterior valley of m3 is, in contrast to S. simplicidens , always present. When both species occur in the same assemblage, S. robustus is always larger than S. simplicidens .

Type locality: Ateca 3, Calatayud–Montalbán basin, Spain (local zone Z, MN 3 , Lower Miocene) .

Stratigraphical distribution: Local zones Z to C (MN 3–MN 4, Lower Miocene).

Geographical distribution: Spain, Portugal, and France.

Figures

Figure 5. Simplomys robustus (De Bruijn, 1967) from San Roque 4A (collection MNCN). A, P4 SR4A-895. B, M1 SR4A-754. C, M2 SR4A-778. D, M3 SR4A-787. E, p4 SR4A-921. F, m1 SR4A-801. G, m2 SR4A-804. H, m3 SR4A-918. Right side specimens underlined.