Pseudepipona aldrichi ( Fox, 1892 ) Fateryga, 2022

Fateryga, Alexander V., 2022, Revision of the Pseudepipona herrichii-group of the eumenine wasps (Hymenoptera: Vespidae: Eumeninae) with the description of two new species from China and Russia, Zootaxa 5154 (2), pp. 101-126 : 104-107

Pseudepipona aldrichi ( Fox, 1892) , stat. nov.

( Figs 1–9 , 25 )

Odynerus aldrichi Fox, 1892: 197–198 , ♀ ♂; type locality: “Brookings, S. Dakota ”, “ Colorado ” [ USA].

Odynerus aldrichii: de Dalla Torre, 1894: 51 , unjustified emendation.

Pseudepipona herrichii aldrichi: Bohart, 1951: 887 .

Pseudepipona variegata mongolica Giordani Soika, 1970b: 330–331 , ♀ ♂; type locality: “Mongolei: Central Aimak , Borulčin Tala, 1400 m, 100 km SO von Ulan-Baator ” [ Mongolia], syn. nov .

Material examined. RUSSIA. Volgograd Province: Sarepta , 1 ♂ [ ZISP] . Omsk Province: Steppe 100 versts S of Omsk, Akmola Province, 23.VIII.1899, 1 ♀, leg. Ignatov [ ZISP]; Vicinity of Omsk, V .1912, 1 ♀, leg. P. Ryabov [ ZISP]; ibid., 15.VII.1912, 1 ♀, leg. N. Vydrina [ ZISP] . Altai Republic: 14 km SE Aktash, Chuya River , 50°13.984′N, 87°45.471′E, 1450 m, 1.VII.2016, 1 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; ibid., 2.VII.2016, 1 ♀, 1 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; ibid., 3.VII.2016, 1 ♀, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CAFK]; ibid., 3.VII.2016, 1 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; ibid., 13.VII.2016, 1 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; Kosh-Agach District , 15 km SE Kuray, Chuya River , 50°11.163′N, 88°07.067′E, 1600 m, 5.VII.2016, 1 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; ibid., 11.VII.2016, 1 ♀, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; Kosh-Agach District , 2 km SE Chagan-Uzun, Balkhash River , 50°05.591′N, 88°23.361′E, 1740 m, 9.VII.2016, 1 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; Kosh-Agach District , 5 km SE Chagan-Uzun, Tydtuyaryk River , 50°04.367′N, 88°25.193′E, 1780 m, 11.VII.2016, 1 ♀, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CAFK]; ibid., 11.VII.2016, 1 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS]; ibid., 8.VII.2019, 1 ♀, leg. Yu.N. Danilov [ CAFK]; ibid., 9.VII.2019, 1 ♀, leg. A. V . Fateryga [ CAFK]; ibid., 10.VII.2019, 1 ♂, leg. A. V . Fateryga [ CAFK]; Onguday, 29. V .1908, 1 ♀, leg. A. Yakobson [ ZISP]; ibid., 3 ♂, leg. W. Steinfeld [ ZISP]; Ulaganskiy District , 12 km SE Aktash, Chuya River , 50°13.852′N, 87°42.981′E, 1570 m, 4.VII.2016, 2 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CAFK]; ibid., 4.VII.2016, 3 ♂, leg. V . M. Loktionov & M. Yu. Proshchalykin [ CFUS] . Khakassia: 25 km N Shira, Tus Lake , 6.VII.2014, 1 ♀, leg. A.S. Lelej, M.Yu. Proshchalykin & V . M. Loktionov [ CAFK]; Altayskiy District , 40 km SE Bely Yar, 3–5 km E Novorossiyskoye, Berezovskoye Lake , 350–380 m, 23. VI .1990, 1 ♀, leg. D. V . Logunov [ ZMMU] ( Fateryga & Mokrousov 2019, as Pseudepipona herrichii ); Shira Lake near Krasnoyarsk, 1 ♂ [ ZISP]; Vicinity of Minusinsk, Shira , 1 ♂, leg. A. Yakobson [ ZISP] . Tyva Republic: Erzin District, vicinity of Erzin , 1000–1100 m, 14–15.VIII.1989, 1 ♂, leg. D. V . Logunov [ FSCV] ( Fateryga et al. 2020, as P. herrichii ) . Krasnoyarsk Territory : 10 km NW Minusinsk, Bystraya River valley, 9.VII.2014, 2 ♂, leg. A.S. Lelej, M.Yu. Proshchalykin & V.M. Loktionov [ CAFK] ; Krasnojarsk [= Krasnoyarsk], 2 ♀, 1 ♂ [ ZISP]; Vicinity of Minusinsk , 30. VI.1924, 1 ♀, leg. N. Filipyev [ ZISP]; Vicinity of Minusinsk, Malaya Minusa , 8.VII.2014, 2 ♂, leg. A.S. Lelej, M.Yu. Proshchalykin & V. M. Loktionov [ CAFK]; Yeniseyskaya Governement, Minusinsk , 11. VI.1897, 1 ♂, leg. Yu. Vagner [ ZISP]; Yeniseyskaya Governement, Minusinsk Uezd, Novoselovo , 10.VIII.1924, 1 ♂ [ ZISP] . Irkutsk Province: Angarsk, 9.IX.1983, 1 ♀, leg. P.G. Nemkov [ FSCV]; Baykalsk , 20.VII.1984, 1 ♀, leg. P.G. Nemkov [ FSCV]; Irkutsk, 7.IX.1983, 1 ♀, leg. P.G. Nemkov [ FSCV]; ibid., 4 ♂, leg. V . E. Yakovlev [ ZISP]; Kirensk, confluence of Kirenka and Lena rivers, 3.VII.1904, 1 ♀, leg. Braudo [ ZISP]; Kultuk , 1 ♀, leg. V . E. Yakovlev [ ZISP]; Nizhneudinsk, 6.VII.1984, 1 ♀, leg. P.G. Nemkov [ FSCV]; Vicinity of Irkutsk, Melnikovo , 1.VII.1912, 1 ♀, leg. N. Sidorov [ ZISP] . Buryatia: 25 km W Kyakhta, Naushki , 2.VIII.1984, 1 ♀, leg. A.S. Lelej [ FSCV] ; 5 km N Naushki, Kharankhoy, 3.VIII.1977, 1 ♂, leg. A.S. Lelej [ FSCV]; Baikal Lake, S coast, Boyarskiy, 11.VIII.1984, 2 ♀, leg. A.S. Lelej [ FSCV]; Kyakhta , 28.VII.1977, 1 ♀, leg. A.S. Lelej [ FSCV]; ibid., 30.VII.1977, 1 ♀, leg. P.A. Lehr [ FSCV]; UstKiran near Kyakhta , 1 ♂, leg. V . E. Yakovlev [ ZISP]. Yakutia: 8 versts W Namtsy, 18.VII.1926, 1 ♀, leg. L. Bianki [ ZISP]; Kempendyay River mouth, inflow of Vilyuy, 9.VII.1927, 1 ♂ [ ZISP]; Lena River , 200 km above Yakutsk, Kharyyalakh Island near Yedey, 5.VII.1985, 1 ♂, leg. Vinokurov [ FSCV]; Markha near Yakutsk, 4.VII.1900, 1 ♂, leg. Olenin [ ZISP]; Modutskiy Nasleg, Namtsy Ulus, VII.1902, 1 ♂, leg. Olenin [ ZISP]; Namtsy Ulus, Aly, 3.VIII.1926, 5 ♀, leg. A. Ivanov [ ZISP]; Near Ulakhan-An River, 150 versts above Yakutsk, 4.IX.1926, 1 ♂, leg. A. Ivanov [ ZISP]; Pokrovsk, 20.VII.1986, 1 ♂, leg. Makarin [ FSCV]; Vilyuy River , Nyurba, 16.VII.1986, 1 ♀, leg. Kaymuk [ FSCV]; Yakutsk , 6.VII.1927, 1 ♂ [ ZISP]; ibid., 11.VII.1927, 1 ♂, leg. Moskvin [ ZISP]; ibid., 16.VII.1927, 2 ♂, leg. Moskvin [ ZISP]; ibid., 17.VII.1927, 2 ♂ [ ZISP]; ibid., 17–21.VII.1927, 1 ♀, 2 ♂, leg. Moskvin [ ZISP]; ibid., 22–23.VII.1927, 1 ♂, leg. Moskvin [ ZISP]; ibid., 24.VII–1.VIII.1927, 2 ♂, leg. Moskvin [ ZISP]; Zhemkonskiy Nasleg, 13.VII.1925, 1 ♂, leg. L. Bianki [ ZISP]. Zabaykalskiy Territory : 15 km W Borzya , 8.VII.1970, 7 ♀, 9 ♂, leg. A.S. Lelej [ FSCV]; ibid., 9.VII.1970, 1 ♂, leg. A.S. Lelej [ FSCV] ; 30 km SW Borzya , 21.VII.1977, 1 ♀, leg. A.S. Lelej [ FSCV] ; 36 km ESE Zabaykalsk, Abagaytuy , 19.VII.1984, 1 ♀, leg. A.S. Lelej [ FSCV] ; 42 km SW Borzya, Durbachi, 23.VII.1984, 1 ♀, leg. A.S. Lelej [ FSCV]; Aleksandrovkiy Plant, Gazimur River, 18.VII.1977, 5 ♂, leg. A.S. Lelej [ FSCV]; Borkha River, upper reaches of Onon, 16.VII.1894, 2 ♂, leg. Kashkar [ ZISP]; Borzya , Chandan, 8.VII.1977, 1 ♂, leg. P.A. Lehr [ FSCV]; Chita, 6.VIII.1912, 1 ♀, leg. A. Kuskov [ ZISP]; Ivan Lake , 45 versts from Chita, 10.VII.1925, 1 ♂, leg. Vinogradov [ ZISP]; Nerchinsk District , 1 ♂, leg. V . Podgorbinskiy [ ZISP]; Nerchinsk Uezd , Soktuy, 17. VI .1925, 1 ♂, leg. Vinogradov [ ZISP]; ibid., 18. VI .1925, 4 ♂, leg. Vinogradov [ ZISP]; ibid., 20. VI .1925, 1 ♂, leg. Vinogradov [ ZISP]; ibid., 23. VI .1925, 1 ♀, leg. Vinogradov [ ZISP]; ibid., 26. VI .1925, 2 ♂, leg. Vinogradov [ ZISP]; Nertschinsk [= Nerchinsk], 1 ♂ [ ZISP]; Unga [Petrovsk-Zabaykalskiy District], 24. VI .1913, 1 ♀, leg. A. Pisarevskiy [ ZISP]; Vicinity of Chalbushi, Shirka River , 8.VII.1970, 1 ♀, leg. N. V . Kurzenko [ FSCV]; Vicinity of Nerchinsk , 1 ♂, leg. V . Podgorbinskiy [ ZISP]; Vicinity of Yedineniye, Onon River, 22.VII.1970, 1 ♀, leg. N. Kurzenko [ FSCV]; Vicinity of Zyulzya , 14.VII.1970, 1 ♂, leg. N. V . Kurzenko [ FSCV]; Vicinity of Zyulzya, left bank of Nercha River , 15.VII.1970, 1 ♀, 1 ♂, leg. N. V . Kurzenko [ FSCV]; ibid., 16.VII.1970, 1 ♀, leg. N. V . Kurzenko [ FSCV] . Amur Province: Tarakon River near Saskal , 13.VIII.1932, 1 ♀, leg. P.A. Lehr [ FSCV] . AZERBAIJAN: Talysh, Lyulyakeran, 15. V .1909, 1 ♀, leg. A. Kiritchenko [ ZISP] . KYRGYZSTAN: Inylchek, 27. VI.1973, 2 ♂, leg. L. Pek [ FSCV]; Kavak Range, Kyzyl Suyek Landmark, 30.VII.1974, 1 ♂, leg. Yu. Tarbinskiy [ FSCV]; Moldo-Too Range, Shavyr Landmark, 20.VIII.1972, 1 ♀, leg. Yu. Tarbinskiy [ FSCV]; Moldo- Too, Shavyr, 20.VIII.1972, 1 ♀, leg. L. Pek [ FSCV]; Onarcha River valley , 10.VIII.1971, 1 ♀, leg. Yu. Tarbinskiy [ FSCV]; Tien-Shan, Sary-Dzhaz, Inylchek, 24.VIII.1974, 3 ♀, leg. L. Pek [ FSCV]; ibid., 26.VIII.1974, 2 ♀, leg. L. Pek [ FSCV]; Tien-Shan, Sary-Dzhaz, Kaindy, 24.VIII.1974, 1 ♀, leg. L. Pek [ FSCV] . KAZAKHSTAN. Akmola Province: Borovoe [currently Burabay], 5.VII.1932, 1 ♀, leg. V . B. Popov [ ZISP]; ibid., 9.VII.1932, 1 ♂, leg. V . B. Popov [ ZISP]; ibid., 17.VII.1932, 1 ♂, leg. V . B. Popov [ ZISP]; ibid., 22.VII.1932, 1 ♀, leg. V . B. Popov [ ZISP]; ibid., 24.VII.1932, 1 ♀, 1 ♂, leg. V . B. Popov [ ZISP]; ibid., 26.VII.1932, 4 ♀, 1 ♂, leg. V . B. Popov [ ZISP]; ibid., 30.VII.1932, 1 ♂, leg. V . B. Popov [ ZISP]; ibid., VIII.1926, 1 ♀, leg. Bey-Bienko [ ZISP]; Kokshetau Uezd, Sarypyldak Lake, VIII.1909, 1 ♀, leg. Piotrovskiy [ ZISP]; Shchuchinsk , 1 ♂ [ ZISP] . East Kazakhstan Province: 18 km SE Georgiyevka [currently Kolbatau], 17. VI .1979, 1 ♂, leg. V . L. Kazenas [ FSCV]; Chagan-Obo , Saur, 20– 28. VI .1910, 1 ♂, leg. B. Karavayev [ ZISP]; Ridder, Tarkhanka, 23.VII.1930, 1 ♀, leg. F. Lukyanovich [ ZISP]; Saur Range , 35 km E Zaysan , 10.VII.1980, 1 ♀, leg. V . L. Kazenas [ FSCV]; Semipalatinsk [currently Semey], 6 ♀, 1 ♂ [ ZISP]; Us-Bulak, Saur , 17.VII.1910, 1 ♂, leg. B. Karavayev [ ZISP] . North Kazakhstan Province: 27 vests behind Petropavlovsk, 12. VI .1894, 1 ♂, leg. N.N. Shiryayev [ ZISP] . Pavlodar Province: Slavgorod Uezd, Severnoye , 6– 7.VII.1922, 1 ♀, leg. A. Reichardt [ ZISP] . MONGOLIA: Bin-Gol [= Bayan-Gol] River valley, SW Kentey, 31.VII– 2.VIII.1924, 1 ♀, leg. P.K. Kozlov [ ZISP]; ibid., 2–3.VIII.1924, 1 ♀, leg. P.K. Kozlov [ ZISP]; Central (Töv) Province, N slope of Bogdoula near Ulaanbaator , 29. VI .1967, 1 ♀, leg. Zaytsev [ ZISP]; Central (Töv) Province, vicinity of Songino , to SW of Ulaanbaator, 1.VII.1967, 1 ♀, leg. Zaytsev [ ZISP]; Dunda-Daydlyga River to E of Urga, 1.VIII.1894, 1 ♂, leg. Kashkar [ ZISP]; Headspring of Tola [= Tuula] River to E of Urga, 3.VIII.1894, 1 ♂, leg. Kashkar [ ZISP]; Khalkha, Murgulin Landmark, 25.VII.1909, 2 ♀, 1 ♂, leg. P.K. Kozlov [ ZISP]; Khalkha , Mzdorgolo Landmark—Khairkhan , 23.VII.1909, 1 ♀, leg. P.K. Kozlov [ ZISP]; Khalkha , Sharkhay-Khunduy Valley , 14.VII.1909, 5 ♂, leg. P.K. Kozlov [ ZISP] ; Övörkhangai Province, from Khan-Khugshin-Ula to Arbay-Khere, 16.VII.1949, 1 ♂, leg. Eglon [ ZISP]; Sangin, Tola [= Tuul] River valley , Khalkha , 25. VI –10.VII.1925, 2 ♂, leg. P. K. Kozlov [ ZISP]; Sangin , Urga, 28.VII.1905, 1 ♀, leg. P.K. Kozlov [ ZISP]; ibid., 29.VII.1905, 2 ♂, leg. P.K. Kozlov [ ZISP]; ibid., 30.VII.1905, 1 ♂, leg. P.K. Kozlov [ ZISP]; Tevshrulekh , 19.VII.1972, 1 ♂, leg. L.N. Medvedev [ FSCV]; Tevshrulekh, Khokhsum, 12. VI .1972, 1 ♂, leg. L.N. Medvedev [ FSCV]; Tola [= Tuul] River , Urga, 1– 3.VII.1905, 1 ♂, leg. P.K. Kozlov [ ZISP]; ibid., 3–5.VII.1905, 1 ♂, leg. P.K. Kozlov [ ZISP]; ibid., 4.VII.1905, 6 ♀, 1 ♂, leg. P.K. Kozlov [ ZISP]; ibid., 5.VII.1905, 9 ♀, 1 ♂, leg. P.K. Kozlov [ ZISP]; ibid., 5–7.VII.1905, 4 ♂, leg. P.K. Kozlov [ ZISP]; ibid., 23.VII.1905, 1 ♂, leg. P.K. Kozlov [ ZISP]; Vicinity of Urga, 23–26. VI .1905, 1 ♀, leg. P.K. Kozlov [ ZISP]; ibid., 7–9.VII.1909, 1 ♂, leg. P.K. Kozlov [ ZISP] . CHINA. Inner Mongolia: Syelchzhy [= Hilinguole] River, Grand Khingan , 27.VIII.1899, 1 ♂, leg. Soldatov [ ZISP]; W slope of Alashan Range, Tszosto, 26. V .1908, 1 ♀, 1 ♂, leg. P.K. Kozlov [ ZISP] . Sichuan: River Fubyanko, Shigaitz—Shintyan , 1.VIII.1893, 1 ♂, leg. G.N. Potanin [ ZISP] . USA. Colorado: Larimer County, Hewlett Canyon, 6000–6500′, 7.VII.1979, 1 ♀, 1 ♂, leg. J.M. Carpenter [ FSCV]; Rifle , 2.VII.1905, 1 ♀, leg. S.A. Rohwer [ FSCV]; ibid., 3.VII.1908, 1 ♂, leg. S.A. Rohwer [ FSCV] .

Observations on iNaturalist. MONGOLIA: Alag-Erdene , 50.50°N, 100.16°E, 7.VIII.2016, 1 ♂, A. Taeger (https://www.inaturalist.org/observations/98791015) . USA. Colorado: [on Geraniaceae ], VI.2020, 1 ♂ (https:// www.inaturalist.org/observations/51092999) ; Boulder County, 40.10°N, 105.39°W [at nest], 28.VII.2015, 1 ♀, P. Wickey (https://www.inaturalist.org/observations/32810598) ; Conifer Drive, South Fork , 37.67°N, 106.65°W, 9.VII.2017, 1 ♀, J.E. Lopez (https://www.inaturalist.org/observations/6987330) ; Las Animas County, 37.09°N, 104.47°W, 27. VI.2020, 1 ♀, H. Eaton (https://www.inaturalist.org/observations/51193028) ; Rocky Mountain National Park , 40.43°N, 105.52°W [on Cirsium sp. ], 28.VIII.2020, 1 ♀, M. McMasters (https://www.inaturalist. org/observations/59366381) ; San Carlos Trail, San Isabel National Forest , Pueblo County, 38.01°N, 105.01°W [on Penstemon glaber (damages corolla robbing nectar)], 28. VI.2014, 1 ♀, R. Bunn (https://www.inaturalist. org/observations/15864003) ; Trinidad , 37.17°N, 104.50°W, 1 ♀, P. Piombino (https://www.inaturalist.org/observations/93515402) ; Wellington , 40.98°N, 105.08°W, 17. VI.2021, 1 ♀, M. Webb (https://www.inaturalist.org/ observations/83564434) . New Mexico: Taos County, 36.14°N, 105.53°W [on Asteraceae ], 15.VII.2017, 1 ♀, R. Pfau (https://www.inaturalist.org/observations/7300707) ; Upper Panchuela Rd. , San Miguel County, 35.83°N, 105.67°W [on Asteraceae ], 21.VII.2021, 1 ♀, V. Remsen (https://www.inaturalist.org/observations/89004911) . Utah: Dixie National Forest, Escalante , 38.00°N, 111.41°W, 25.VII.2020, 1 ♀ (https://www.inaturalist.org/observations/55991064) . Wyoming: Teton County, 43.63°N, 110.81°W, 19.VII.2017, 1 ♀, S. Whipple (https://www.inaturalist.org/observations/9567621) ; Yellowstone National Park , 44.43°N, 110.59°W [on Asteraceae ], 12.VIII.2015, 1 ♀, T. C. Jones (https://www.inaturalist.org/observations/65466212) . CANADA. Alberta: Fish Creek Provincial Park, Calgary , 50.92°N, 114.00°W, 1.VII.2020, 1 ♀, M. Summers (https://www.inaturalist.org/observations/52853738) ; Inglewood, Calgary , 51.26°N, 114.01°W, 16. VI.2020, 1 ♀ (https://www.inaturalist.org/observations/51925495) . British Columbia: Central Kootenay , 49.32°N, 117.65°W, 10. VI.2021, 1 ♂ (https://www.inaturalist.org/observations/82609020) ; Sunglo Drive, Penticton , 49.49°N, 119.63°W [on Rosaceae ], 29. V.2021, 1 ♂ (https://www. inaturalist.org/observations/80803661) . Saskatchewan: Division No. 4, 49.15°N, 107.51°W [on Eriogonum sp. ], 9. VI.2021, 1 ♂, S. Simpkins (https://www.inaturalist.org/observations/83214534) ; Division No. 4 [on Fabaceae ], VII.2020, 1 ♂, T. Miller (https://www.inaturalist.org/observations/56470132) ; Grasslands National Park, Val Marie, 49.20°N, 107.66°W [on Penstemon sp. (robbing nectar)], 23. VI.2021, 1 ♂, E. Gorda (https://www.inaturalist. org/observations/92558993) ; Val Marie No. 17, 49.18°N, 107.70°W, 7. VI.2021, 1 ♂, S. Simpkins (https://www. inaturalist.org/observations/104236676) ; ibid., 9. VI.2021, 1 ♂, S. Simpkins (https://www.inaturalist.org/observations/83018848) .

Additional literature records. MONGOLIA: Chentej Aimak, 7 km NO von Somon Mörön, 1200 m ( Giordani Soika 1970b). USA. Alaska: Denver Glacier, Skagway ( Bohart 1939). Colorado: Estes Park ( Bohart 1939). Idaho ( Krombein 1979). Montana: Silver Bow County ( Bohart 1939). South Dakota: Brookings ( Fox 1892). Washington: Grand Coulee ( Bohart 1939). Wyoming: Grand Teton National Park ( Bohart 1939); Grand Teton National Park, about 4 km WNW of the Moran Post Office ( Evans 1978, as Pseudepipona herrichii ); Jackson Hole ( Evans 1970).

Diagnosis. Females of this species can be distinguished from other members of the P. herrichii -group by the following combination of characters: clypeus 1.05–1.10× as wide as long, entirely black or with a basal whitish band; scutellum with a thin apical band equal to that on the metanotum; propodeum with a weakly developed transverse carina between the shelf and concavity forming laterally blunt angles; propodeal valvula rounded posteriorly; T1 without three small longitudinal carinae. In the males, the clypeus is 1.10–1.15× as wide as long; cuspis with a very broad dorsal process which is much shorter than its width at the base; base of the digitus attached directly to the top of the cuspis apex; aedeagus with a median expansion; ventral lobe of the aedeagus large, in lateral view much larger than the median expansion. In both females and males from the Palaearctic region, the tegula and often the band on the pronotum are reddish; this character is useful to distinguish Asiatic specimens of P. aldrichi from the similar P. herrichii .

Distribution. Russia (Volgograd Province, Omsk Province, Altai Republic, Khakassia, Tyva Republic, Krasnoyarsk Territory, Irkutsk Province, Buryatia, Yakutia, Zabaykalskiy Territory, Amur Province), Azerbaijan, Kyrgyzstan, Kazakhstan (Akmola Province, East Kazakhstan Province, North Kazakhstan Province, Pavlodar Province), Mongolia, China (Inner Mongolia, Sichuan), USA (Alaska, Colorado, Idaho, Montana, New Mexico, South Dakota, Utah, Washington, Wyoming), Canada (Alberta, British Columbia, Saskatchewan) ( Fig. 25 ).

Remarks. No significant differences between Asiatic ( P. herrichii mongolica ) and North American ( P. herrichii aldrichi ) forms of this species were revealed in the male genitalia. The only difference between them is the coloration of the tegula (reddish in the Asiatic specimens and usually whitish in the North American ones) and the light band on the pronotum (usually reddish in the Asiatic specimens and whitish in the North American ones). Bohart (1939) have already pointed out that P. aldrichi is present in China. I agree with him that this species has an Asiatic origin and it was distributed into North America by the way of Alaska.

Bionomics. Two nests of this species were briefly described by Evans (1978) in USA. They were excavated closely to each other, by two females, in the hard sandy loam of a path. Apparently, the females used water to moisten the substrate during the excavation but that was not pointed out. A female was observed carrying earthen pellets from the nest burrow by flight and dropping them in 0.6–0.9 m from the nest. Nests burrows were about 9.5 in depth; neither nest had a turret at the entrance. The number of cells in the nests was two and one; the two-celled nest had a branched structure. Provision was caterpillars of a gelechiid moth ( Lepidoptera : Gelechiidae ); up to 17 prey items were recorded in a cell. Adult feeding was recorded in USA and Canada on flowers of various plants in the families Asteraceae , Fabaceae , Geraniaceae , Rosaceae , and Polygonaceae . Both females and males were also observed robbing nectar from flowers of Penstemon spp. (Plantaginaceae) (see Observations on iNaturalist).

Figures

FIGURES 1–6. Pseudepipona aldrichi (Fox, 1892). 1, 3, 5, habitus in dorsal view (1, 3, ♀; 5, ♂); 2, 4, 6, head in frontal view (2, 4, ♀; 6, ♂). 1, 2, Russia: Altai Republic; 3, 4, USA: Colorado; 5, 6, Russia: Zabaykalskiy Territory. Scale bars 1 mm.

FIGURES 7–24. Male genitalia of Pseudepipona spp. 7–9, P. aldrichi (Fox, 1892); 10–12, P. herrichii (de Saussure, 1856); 13–15, P. kozhevnikovi (Kostylev, 1927); 16–18, P. tingjingliae Fateryga, sp. nov.; 19–21, P. tricarinata (Kokujev, 1913); 22–24, P. valentinae Fateryga, sp. nov. 7, 10, 13, 16, 19, 22, left paramere and volsella in lateral view from inside (bd, base of digitus; ca, cuspis apex; dp, dorsal process of cuspis); 8, 11, 14, 17, 20, 23, aedeagus in dorsal view (me, median expansion); 9, 12, 15, 18, 21, 24, aedeagus in lateral view (vl, ventral lobe). Scale bar 1 mm.

FIGURES 25–28. Distribution of Pseudepipona spp. 25, P. aldrichi (Fox, 1892); 26, P. herrichii (de Saussure, 1856); 27, P. kozhevnikovi (Kostylev, 1927); 28, P. tingjingliae Fateryga, sp. nov. (black dots), P. tricarinata (Kokujev, 1913) (white dots), and P. valentinae Fateryga, sp. nov. (grey dots).