Arenigena marcida (Coquillett)

Webb, Donald W., 2009, A revision of the genera Acrosathe Irwin and Lyneborg, Arenigena Irwin and Lyneborg, and Litolinga Irwin and Lyneborg (Diptera: Therevidae: Therevinae) from the Nearctic Region, Zootaxa 2091 (1), pp. 1-67 : 46-49

Arenigena marcida (Coquillett)

(Figs. 80–87, 96)

marcida Coquillett (1893b:224 key, 228 male desc.), Aldrich (1905:247 cat.), Kröber (1912: 222 male key, 229 spec. list;

1913:32 dist., 38 male key; 1914: 38 male key, 44 desc.), Cole (1923a:69 desc., 70 dist., Figs. 56 fem. frons, 79 ant.,

97 male genit.; 1965:351 cat.) ( Psilocephala ); Irwin and Lyneborg (1981a:240 comb. change, spec. list) ( Arenigena ). Type locality USA, California, Los Angeles County. Lectoype male in USNM. brunnea Kröber (1914:46 fem. desc.), Cole (1923a:76 desc.; 1965:351 cat.) ( Psilocephala ); Irwin and Lyneborg

( 1981a:240 comb. change, spec. list) ( Arenigena ). Type locality USA, Arizona, Oracle. Holotype female in USNM

(Type No. 26020). New synonym.

The holotype female of Psilocephala brunnea Kröber was compared with females of Arenigena marcida and found to be conspecific based on the characteristics of the female furca.

Diagnosis. Arenigena marcida is similar to A. bajaensis in having the inner gonocoxal process (Fig. 82) narrow, elongate and the distiphallus (Fig. 86) thick basally and broadly curved posteroventrally. It differs from A. bajaensis in the male frons lacking setae; the female frons having short, reddish brown setae; the distiphallus (Fig. 84) broadly expanded laterally; and the furca (Fig. 87) with the anterior margin narrow.

Description of MALE. Variation (n=10).

Body length 7.0–9.4, 8.0 mm.

FIGURES 80–87. Arenigena marcida . 80. Tergite 8, epandrium, cerci. 81. Sternite 8, gonocoxite ventral view. 82. Gonocoxite, lateral view. 83. Gonostylus lateral view. 84. Aedeagus, dorsal view. 85. Aedeagus, ventral view. 86. Aedeagus, lateral view. 87. Female internal reproductive organs. Scale = 0.1 mm.

Head. Length 0.84–1.16, 0.95 mm. Ocellar tubercle dark reddish brown, pubescence gray, dense; setae dark reddish brown, short. Frons pubescence white to pale gold, dense; setae absent. Antenna yellow, second and third flagellomeres dark brown; antenna/head length 0.54–0.78, 0.58; scape length 0.16–0.22, 0.19 mm, width 0.12–0.18, 0.14 mm, length/width 1.1–1.6, 1.3, scape/pedicel length 1.6–3.1, 2.4, scape/flagellum width 0.7–1.0, 0.8, setae pale yellow to yellowish brown, short; pedicel length 0.07–0.10, 0.08 mm, width 0.12–0.16, 0.14 mm, length/width 0.4–0.7, 0.6; flagellum (narrow apical portion of first flagellomere short), length 0.30–0.44, 0.37 mm, width 0.16–0.22, 0.14 mm, length/width 1.7–2.7, 2.0, flagellum/scape length 1.6–2.3, 2.0. Parafacial pubescence yellowish gray. Gena pubescence whitish gray; setae white, elongate. Maxillary palpus yellow; length 0.32–0.48, 0.40 mm, width 0.06 mm, length/width 5.3–8.0, 6.7; setae white to pale yellow, elongate. Occipital setae white, elongate white, abundant dorsally.

Thorax. Macrosetae 3 np, 2 sa, 1–2, dc. Mesonotum pubescence gray, dense; setae white, elongate. Pleuron pubescence gray, dense; setae white, elongate on propleuron, proepimeron (occasionally absent), anepisternum, katepisternum (sparse, variable, in anterodorsal and anteroventral patch, in mediodorsal patch, or in narrow band along anterior margin), laterotergite, metanepisternum, and scutellum. Wing. Membrane pale whitish gray, veins and cross-veins brown with brown spot at fork of R 4+5, r-m, apex of discal cell, apex of cells bm and cup; length 5.3–7.0, 6.2 mm, width 2.0–2.5, 2.2 mm, length/width 2.3–3.1, 2.7; pterostigma dark brown. Legs. Coxae pubescence white, dense; setae white, elongate. Femora dark reddish brown, pubescence white; ventral setae white, elongate, dorsal setae white, appressed. Tibiae yellow. Tarsi 1 dark yellow; 2–5 brown.

Abdomen. Pubescence white, dense; dorsal setae white, elongate, erect, abundant; lateral setae white, elongate. Terminalia (MEI 113211) brown. Tergite 8 (Fig. 80), setae pale yellow. Sternite 8 (Fig. 81), rectangular, sides rounded, posterior margin slightly emarginate; setae pale yellow, separated medially on posterior margin. Epandrium (Fig. 80), posterolateral margin extended, pointed; setae yellow. Hypoproct (Fig. 80), posterior margin slightly emarginate. Gonocoxite ventral view (Fig. 81), posterior margin rounded, setae yellow, macrosetae absent; lateral view (Fig. 82), posterior margin rounded, inner gonocoxal process narrow, elongate. Gonostylus lateral view (Fig. 83), sinuate. Aedeagus with dorsal apodeme (Fig. 84) broadly rounded laterally, anterior margin emarginate; ventral apodeme (Fig. 85), broad; distiphallus dorsal view (Fig. 83) broad basally, tapered posteriorly; lateral view (Fig. 86) thick basally, extending posterior in broad curve; ejaculatory apodeme (Figs. (84–85) not reaching anterior margin of dorsal apodeme.

FEMALE. Similar to male except for following. Variation (n=10).

Body length 7.8–10.4, 9.4 mm.

Head. Length 0.96–1.20, 1.07 mm. Frons pubescence yellowish gray; setae reddish brown, short, scattered. Antenna/head length 0.59–0.70, 0.63; scape length 0.16–0.24, 0.21 mm, width 0.14–0.18, 0.17 mm, length/width 0.9–1.6, 1.3, scape/pedicel length 1.6–3.0, 2.5, scape/flagellum width 0.7–1.0, 0.8; pedicel length 0.08–0.12, 0.09 mm, width 0.14–0.18, 0.16 mm, length/width 0.5–0.6, 0.5; flagellum length 0.34–0.44, 0.39 mm, width 0.16–0.24, 0.21 mm, length/width 1.7–2.5, 1.9, flagellum/scape length 1.7–2.1, 1.9. Maxillary palpus length 0.34–0.52, 0.43 mm, width 0.06–0.08, 0.07 mm, length/width 5.0–7.7, 6.0.

Thorax. Macrosetae 3 np, 2 sa, 1 dc. Wing. Length 6.3–7.8, 6.8 mm, width 2.2–3.0, 2.5 mm, length/width 2.5–3.3, 2.8.

Abdomen. Pale orange brown, pubescence light whitish gray; dorsal setae on tergites 1–4 white, appressed, tergites 5–8 dark reddish brown. Terminalia (MEI 109201) reddish brown. Furca (Fig. 87) quadrate; length 0.50 mm, width 0.34 mm; anterolateral projection short.

Immature stages. Unknown.

Distribution. Arenigena marcida is known from Mexico (Baja California Norte, Baja California Sur, Sinaloa) and the United States (Arizona, California, Nevada, New Mexico, Texas) ( Fig. 97 ).

Habitats and Phenology. Arenigena marcida has been hand-netted or sweep-netted in sandy creek bottoms and collected in pit traps, Malaise traps (in coastal sage scrub), ethylene glycol intercept traps, and at mercury vapor and blacklights. Adults have been collected from February through November.

Specimens examined. Type specimens. Two male syntypes of Psilocephala marcida Coquillett were examined. One male is labeled "Los Angeles Co., Cal., Collection Coquillett" USNM Type No. 10415 (MEI 141533) and the other is labeled "San Diego Co. CAL., Collection Coquillett" USNM Type No. 10415 (MEI 141547). The male from Los Angeles County (MEI 141533) is designated as the lectotype. This lectotype is designated in order to fix the concept of Psilocephala marcida Coquillett and to ensure universal and consistent interpretation of the same. The specimen from San Diego County (MEI14147) is designated as a Paralectotype. The holotype female of Psilocephala brunnea Kröber (MEI 112374) is labeled "Oracle, 29.6 Ar, Coll Hubbard & Schwarz" and is in the USNM (Type No. 26020). Other specimens. MEXICO. BAJA CALIFORNIA NORTE. 6.4 km S La Rumorosa [32.491, -116.05], 1321m, 18.IV.1964, Schlinger, E.I., 1♂ MEI 119394 (MEIC), sandy creek bottom; 2.4 km E Santa Rosalillita, Arroyo Santo Dominguito [28.67, -114.208], 25.III.1970, Fisher, E.M., Fisher, J.L., 1♀ MEI 109239 (MEIC); 35.4 km S El Arco [27.71, -113.401], 13.IV.1968, Irwin , M.E., 1♂ MEI 109250 (MEIC); Sierra San Pedro Martir, east base, Diablo Canyon [31.064, -115.361], 671m, 6–12.VI.1997, Irwin , M.E., Schlinger, E.I., 1♀ MEI 099894 (MEIC), Malaise trap at waterhole. BAJA CALIFORNIA SUR. 1.6 km S Punta Colorado, Isla San Jose [24.984, -110.622], 8–9.IV.1974 Doyen, J.T., 1♀ MEI 154600 (EMEC); 30 km E La Ribera, Rancho Las Barracas [23.6, -109.28], 21–24.III.1982 Irwin , M.E., Schlinger, E. I., 1♀ MEI 114534 (MEIC); 49.9 km S El Crucero, 6.XI.1995, Ewart, W., Dickson, R., 1♀ MEI 119143 (MEIC), sweep-net grasses, forbs; El Triunfo [23.786, -110.13], 13.VII.1938, Michelbacher, Ross, 1♂ MEI 119201 (EMEC). SINALOA. 25.8 km S Guamuchil, 16.III.1962, Stange, L.A., 2♂ MEI 119188–9 (MEIC); 22.5 km W Guaymas [27.921, -111.103], 13.XI.1965 Ewart, W., Dickson, R. 3♂ MEI 119182, 119208, 119129 (MEIC), at light at light. UNITED STATES. ARIZONA. Cochise County. 12.9 km NE Portal [31.996, -109.059], 9.V.1956, Statham, M., 1♂ MEI 119180 (AMNH); 27.4 km E. Douglas [31.344, -109.298], 12.VIII.1975, Pinto, J.D., 2♂ MEI 119191, 119193 (UCR), blacklight; Douglas [31.344, -109.545], 7.IX.1969, 1♂ MEI 026923 (AMNH); Portal [31.914, -109.141], 1219–1524m, 16.IX.1958, Weems, C., 1♀ MEI 119127 (FSCA); 3.2 km NE Portal [31.934, -109.12], 29.VIII.1965, Davidson, J.H., Davidson, J.M., Cazier, M.A., 1♂ MEI 119415 (ASUT). Graham County. Whitlock Mountains, Dripping Spring [32.581, -109.384], 5.VIII.1976, Chandler, D.S., 1♂ MEI 078449 (UAIC), blacklight. Maricopa County. 29 km S Gila Bend [32.687, -112.716], 18.IV.1965, Gorodenski, S.A., Davidson, J.M., Cazier, M.A., 1♂ MEI 119385 (ASUT); El Mirage [33.613, -112.32], 17.X.1986, Fritz, R.F., 1♂ MEI 072743 (EMUS); Sun City [33.598, -112.271], 11.X.1986, Fritz, R.F., 1♀ MEI 072744 (EMUS); Thunderbird Regional Mountain Park [33.693, -112.186], 13.IV.1975, Wielgus, R., 1♂ MEI 170258 (ASUT). Navajo County. Kayenta, Laguna Creek [36.717, -110.25], 13.VII.1967, Davidson, J.H., Davidson, J.M., Cazier, M.A., 1♂ MEI 119126 (ASUT). Pima County. 16 km W Tucson [32.24, -111.13], 900m, 10–15.IV.1995, Prchal, S., 1♀ MEI 081416 (LACM), Malaise trap; 8 km E Robles Junction, Avra Valley [32.067, -111.25], 10.IV.1996, Yanega, D., 1♂ MEI 119174 (INHS); 14–24.IV.1996, Yanega, D., 2♂ MEI 119173, 119198, 1♀ 170254 (INHS), blacklight; Baboquivari Mountains, Baboquivari Peak [31.781, - 111.579], 8.V.1938, Parker, F.H., 1♂ MEI 078487 (UAIC); Sabino Canyon [32.328, -110.801], 3.IX.1963, Yesterby, V.L., 1♀ MEI 119136 (MEIC); 2.VI.1964, Gorodenski, S.A., 1♂ MEI 170255, 1♀ 170256 (ASUT); Sahuarita, Santa Cruz River [31.958, -110.955], 19.IV.1957, Butler, G., Werner, F., 1♂ MEI 078495 (UAIC), sweep-net; Santa Catalina Mountains, Sabino Canyon [32.328, -110.801], 853m, 12.III.1961, Painter, E.M., 1♂ MEI 119209 (KSUC); 4.IX.1965, O'Brien, L., O’Brien, C.W., 1♂ MEI 119213 (MEIC); St. Catalina Mountains, W slope of Pusch Peak, near Hardy Road, Highway 80 [32.372, -110.938], 853m, 17.V.1963, Mickel, C.E., 1♀ MEI 078294 (MEIC), blacklight; Tucson [32.222, -110.926], 1954, collector?, 1♂ MEI 078447 (UAIC); 13.VIII.1936, Knull, J.N., 1♀ MEI 114592 (OSU); 6.IX.1938, Crandall, R.H., 1♀ MEI 119130 (AMNH); 14.IV.1940, 3♂ MEI 119181–2, 119195 (AMNH); 14.V.1953, Dietrich, A., Dietrich, H., 1♀ MEI 119232 (CUIC); 21.IV.1960, Sipes, C., 1♂ MEI 119215 (MEIC); 18.VI.1979, Werner, F.G., 1♀ MEI 078486 (UAIC), flight trap; Tucson, N. Nanini [32.222, -110.926], 29.IV.1979, Nutting, W.L., 3♂ MEI 078489, 078493–4, 2♀ 078490, 078499 (UAIC); Tucson, near Oracle [32.228, -110.978], 8.IV.1985, Nutting, W.L., 1♀ MEI 078496 (UAIC); 18.VIII.1985, 1♀ MEI 078497 (UAIC), pool; 20.VI.1990, 1♀ MEI 078476 (UAIC); Organ Pipe Cactus National Monument, [32.042, -112.875], 16.IV.1948, Melander, A.L., 1♂ MEI 113252, 1♀ 113250 (USNM), at light; 22.III.1961, Stehr, F.H., 3♂ MEI 078293, 078295–6, 3♀ 078297–9 (UMSP), at light in campground; 9.IV.1960, Santana, F.J., 1♂ MEI 128576 (MEIC), 7.V.1960, Butler, G., 1♂ MEI 170257 (MEIC), blacklight trap; Caretaker's house [32.224, -111.159], 16.X.1982, 1♂ MEI 118857, 1♀ 118856 (SASI); 11.V.1984, 1♂ MEI 118854 (SASI); 22.V.1984, 1♂ MEI 118853 (SASI); 13.VI.1984, 1♂ MEI 118855, 1♀ 118852 (SASI). Pinal County. near Sacaton [33.079, -111.738], 14.VI.1993, Lindsey, M.L., 1♀ MEI 078488 (UAIC). Santa Cruz County. 1.6 km E Sonoita [31.671, -110.642], 1433m, 15.V.1961, Painter, R.H., Painter, E.M., 1♂ MEI 119210 (KSUC). Yavapai County. Dead Horse Ranch [Dead Horse Ranch State Park], 945m, 4.VI.1979, Rohde, M., 1♀ MEI 078491 (UAIC); Congress [34.16, -112.839], 914m, 15.VI.1974, Rockwood, A., Foster, N., Kolner, M., 1♂ MEI 119184 (MEIC); Yava [34.472, -112.89], 1158m, 17.VIII–7.IX.1995, Beal, R.S., 1♀ MEI 116985 (CSUC), ethylene glycol intercept trap. Yuma County. Kofa Mountains 4.V.1963, Fisher, E., 1♂ MEI 119176 (UCR); Martinez Lake, near McAllister Wash [33.017, -114.483], 27.III.1961, Toschi, C.A., 1♂ MEI 119396 (MEIC). CALIFORNIA. Contra Costa County. 3.2 km E Antioch [38.005, -121.805], 16.V.1967, Opler, P.A., 1♂ MEI 170204 (EMEC); Antioch [38.005, - 121.805], 20.IV.1930, Hurd, P.D., 1♀ MEI 119063 (EMEC); MacNeill, C.D., 1♂ MEI 119384, 1♀ 119380 (MEIC); 15.V.1958, Linsdale, D.D., 1♀ 170201 (MEIC); Powell, J., 3♂ MEI 119089, 119382, 1193873 (EMEC), at light; 14.IV.1977, 11♂ MEI 119066–7, 119078–9, 119084–6, 119381, 119387–9, 2♀ 119091, 119403 (EMEC); 7.V.1978, Johnson, J.B., 1♀ MEI 079929 (WFBM). Fresno County. 29 km SW Mendota, Ciervo Hills [36.569, -120.565], 18.VII.1975, Powell, J., 1♂ MEI 119420 (EMEC); 6.4 km W Coalinga, dry bed Waltham Creek [36.141, -120.42], 28.VIII.1952, 1♂ MEI 156358 (CAS); Coalinga [36.141, -120.363], 17.III.1931, Van Duzee, E.P., 1♀ MEI 119410 (MEIC); Fresno [36.748, -119.771], 24.III.?, Schwarz, E.A., 1♂ MEI 113253 (USNM); 3.IV.1923, Phillips, M.E., 1♀ MEI 113206 (USNM). Imperial County. 12.9 km NW Glamis, Algodones Dunes [33.053, -115.191], 11.IV.1990, Webb, D.W., Irwin , M.E., 1♂ MEI 119408 (INHS); 14.5 km S Walter's Camp [33.109, -114.689], 28.IX.1980, Hardy, A.R., Andrews, F.G., 3♂ MEI 170205, 170207, 170209, 2♀ 170206, 170208 (MEIC), blacklight; 25.8 km NW Westmorland [33.204, - 115.79], 2.XI.1974, Powell, J.A., 2♂ MEI 119140, 119414 (EMEC), UV trap; 3.2 NW Glamis [33.018, - 115.091], 3.XI.1974, Powell, J.A., 1♀ MEI 119413 (EMEC); 4.8 km N Glamis [33.041, -115.071], date?, Washbauer, M., 1♂ MEI 073035 (CSCA), blacklight; 4.8 km NW Glamis [33.028, -115.102], 24.III.1972, Sumlin, W.D., 1♀ MEI 119131 (MEIC); 5.6 km NW Glamis [33.033, -115.107], 13.IV.1968, Bellet, E.M., 1♂ MEI 119158 (MEIC); 9.7 km NE Ogilby, Glamis Dunes [32.879, -114.776], 19.III.1980, Werner, Olson, Metz, MacLachlan, 1♂ MEI 078484 (UAIC). Inyo County. 11.3 km S Oasis [37.383, -117.914], 1539m, 23–24.VI.1992, Irwin , M.E., Yeates, D.K., 1♂ MEI 119424 (MEIC), sweeping sagebrush; 33.8 km E Keeler [36.487, -117.569], 28.VI.1980, Toft, C.A., 1♂ MEI 119417 (UCDC); Antelope Springs [37.331, -118.086], 28.VI.1962, Buckett, J.S., 1♀ MEI 170250 (MEIC); Eureka Valley [37.111, -117.707],?. III.1978, Giuliani, Hardy, Andrews, 1♂ MEI 170221, 1♀ 170222 (CSCA), pitfall trap;?. IV.1978, 1♂ MEI 170220, 1♀ 170224 (CSCA), pitfall trap; Sand Flat [36.655, -117.432], 1341m, 19.IV.1986, Giuliani, D., 1♂ MEI 072553 (CSCA); Owens Valley, 30.V–28.VI.1980, 3♂ MEI 170212–4, 7♀ 170214–9, 170223 (CSCA), pit trap. Kern County. 1.6 km NW Tupman [35.304, -119.361], 19.III.1975, Doyen, J.T., 1♂ MEI 119422 (EMEC); 6.4 km NE Inyokern [35.688, -117.771], 16.IV.1959, Linsdale, D.D., 1♂ MEI 119400 (MEIC); Bakersfield [35.356, - 119.005], 20.VII.1951, Isaak, L.W., 1♂ MEI 119407 (MEIC). Kings County. 3.2 km SW Kettleman City [35.988, -119.981], 30.III.1976, Doyen, J., Rude, P., Morrison, R., 1♂ MEI 119423 (EMEC). Los Angeles County. 1♂ MEI 141533 (USNM);?.IX.?, 4♂ MEI 113204–5, 113207, 113209 (USNM); 9.7 km NE Three Point [34.798, -118.536], 11.IV.1962, Schlinger, E.I., Hall, J.C., 1♂ MEI 119068 (UCR); Angeles National Forest, Islip Canyon [34.2, -117.867], 12.VI.1974, Wilder, D.D., 1♂ MEI 156350 (CAS); Boquet Canyon [34.429, -118.516], 9.VI.1960, Schlinger, E.I., 1♂ MEI 119219 (UCR); Los Angeles [34.052, -118.243], date?, Banks, N., 1♂ MEI 081091 (MCZ); Mint Canyon [34.429, -118.441], 24.VI.1949, Isaak, L.W., 1♂ MEI 119200 (MEIC); Tanbark Flat [34.204, -117.76], 17.VII.1950, Hurd, P.D., 1♀ MEI 119224 (EMEC); 26.VI.1952, Grigarick, A.A., 1♂ MEI 119218 (MEIC); 6.VII.1952, Linsley, J. 1♂ MEI 119199 (EMEC). Marin County. Bass Lake [37.963, -122.751], 7.VI.1940, Cazier, M.A., 1♂ MEI 119183 (AMNH). Merced County. 3.2 km South of Delhi [37.402, -120.78], 3.V.1961, Snelling, R.R., 2♂ MEI 156353, 156355 (CAS). Orange County. Costa Mesa [33.641, -117.919], 12.VI.1961, Irwin , M.E., 1♂ MEI 170211 (MEIC); Laguna Beach [33.561, -117.818], date?, Baker, 1♂ MEI 115635 (USNM); San Juan Canyon, 229m, 11.V.1968, Schlinger, E.I., 1♀ MEI 119144 (UCR); Newport Beach [33.619, -117.928], 6.V.1962, Michelbacher, A.E., 1♂ MEI 119197 (EMEC). Riverside County. 1.6 km N Snow Creek [33.904, -116.683], 25.IX.1965, Dickson, R.C., 1♂ MEI 170225 (MEIC); 6 km N Desert Center [33.766, -115.402], 18.III.1986, Steiner, W.E., Lowry, J.E., 1♂ MEI 079114 (MEIC); 6.4 km S Beaumont, Potrero Creek [33.88, -116.978], 1.VII.1990, Ballmer, G.R., Hawks, D.C., 1♂ MEI 078133 (UCR), at light; 6.4 km S Palm Desert, Deep Canyon, P.L. Boyd Desert Research Center, Marker 5680 [33.665, -116.374], 18.V.1969, Irwin , M.E., 1♀ MEI 119142 (MEIC), Malaise trap; 24.V.1969, Irwin , M.E., Frommer, S.I., UCR 19121 (UCR), Malaise trap; 6.5 km NE Alpine Village, near Cahulla Tewanet Vista Point [33.597, -116.422], 1200m, 2–4.VI.2002, Irwin , M.E., Parker, F.D., 1♂ MEI 170226 (MEIC), Malaise trap; Aguanga [33.447, -116.857], 11.V.1953, Wilcox, J., 1♀ MEI 119225 (EMEC); Big Morongo River, 13.IV.1989, Freidberg, A., 1♀ MEI 119071 (TAUI); Box Springs Mountains, Riverside [33.971, -117.302], 26.VI.1969, Andrews, F.G., Hall, J.C., Irwin , M.E., 3♀ MEI 119154, 119222–3 (UCR), Malaise trap; 4.VII.1969, 1♀ MEI 119233 (UCR), Malaise trap; 16.VII.1969, 2♀ MEI 119230–1 (UCR), Malaise trap; Garner Valley, Morris Ranch Road, Tripp Meadow [33.643, -116.595], 1725m, 4.VI.2002, Irwin , M.E., Parker, F.D., 1♀ MEI 170248 (MEIC), Malaise trap; Menifee Valley [33.65, -117.2], 475m, 15–22.V.2000, Frommer, S.I., 2♂ MEI 139395, 170270, 1♀ 139403 (UCR), Malaise trap, coastal sage scrub; 1–5.VI.2000, 1♂ MEI 137186 (UCR), Malaise trap, coastal sage scrub; 5–12.VI.2000, 1♀ MEI 139404 (UCR), Malaise trap, coastal sage scrub; 19–26.VI.2000, 1♀ MEI 139406 (UCR), Malaise trap, coastal sage scrub; Morris Ranch Road, Golf Flat, Garner Valley [33.643, -116.595], 1725m, 4.VI.2002, Irwin , M.E., Parker, F.D., 1♀ MEI 170249 (MEIC), Malaise trap; Palen Dunes [34.111, -115.624], 27.IV.1978, Andrews, F.G., Hardy, A.R., 1♂ MEI 073037 (CSCA), Hg vapor light; Palm Springs [33.83, -116.544], 16.II.1935, Melander, 1♂ MEI 113251 (USNM); Palm Springs, Smoke Tree Ranch [33.789, -116.529], 6.IV.1949, collector?, 1♂ MEI 037085 (USNM); Pinyon Flat [33.603, -116.455], 7.V.1986, Goodward, D., 1♂ MEI 119177 (UCR); Riverside [33.94, -117.397], 13.IV.1964, Rauch, P.A., 1♀ MEI 119151 (UCR); 24.IV.1966, Schlinger, E.I., 1♀ MEI 119146 (UCR); 1.V.1969, Hall, J.C., 1♀ MEI 119045 (UCR); 12.V.1969, 1♂ MEI 119157 (UCR); 19.V.1969, Andrews, F.G., 1♂ MEI 119156, 4♀ 119237, 119106, 119116, 119141 (UCR), Malaise trap; 21.V.1969, Hall, J.C., 1♀ MEI 119117 (UCR); 24.V.1969, Andrews, F.G., 2♀ MEI 119235–6 (UCR), Malaise trap; 27.V.1969, Hall, J.C., 1♀ MEI 119111 (UCR); 30.V.1969, Irwin , M.E., 2♀ MEI 119145, 119234 (MEIC), Malaise trap; 2.VI.1969, Hall, J.C., 1♀ MEI 119125 (UCR), Malaise trap; 3.VI.1969, 1♀ MEI 119120 (UCR), Malaise trap; 6.VI.1969, Irwin , M.E., 2♀ MEI 119147, 119238 (MEIC), Malaise trap; 9.VI.1969, Hall, J.C., 1♀ MEI 119107 (UCR); 16.VI.1969, 2♀ MEI 119110, 119114 (UCR); Irwin , M.E., Hall, J.C., 1♀ MEI 119150 (MEIC), Malaise trap; 30.VI.1969, Hall, J.C., 1♂ MEI 119166, 1♀ 119122 (UCR), Malaise trap; 10.VII.1969, 1♀ MEI 119121 (UCR), Malaise trap; 15.VIII.1969, 1♀ MEI 119112 (UCR), Malaise trap; 13.IX.1969, 1♂ MEI 119165 (UCR), Malaise trap; 13.IV.1970, 1♀ MEI 170244 (UCR); 2.V.1970, 3♂ MEI 119164, 119168–9, 1♀ 119123 (UCR), Malaise trap; 8.V.1970, 1♀ MEI 119155 (UCR), Malaise trap; 12.V.1970, 3♂ MEI 119159, 119167, 119196 (UCR), Malaise trap; 18.V.1970, 1♀ MEI 119119 (UCR), Malaise trap; 22.V.1978, 1♀ 170247 (UCR); 30.V.1978, 1♀ MEI 119105 (UCR); 9.VI.1978, 1♀ MEI 119104 (UCR); 13.VI.1978, 2♀ MEI 119109, 119118 (UCR); 21.VI.1978, 1♀ MEI 170245 (UCR); 26.VI.1978, 1♀ MEI 119113 (UCR); 17.VII.1978, 1♀ MEI 119108 (UCR); 18.VII.1978, 1♀ MEI 119103 (UCR); Riverside, 131 Sir Damas Drive [33.991, -117.346], 17.V.1976, Stone, M.W., 1♂ MEI 119194 (UCR); 16.VI.1976, 1♀ MEI 170252 (UCR); 18.VII.1977, 1♂ MEI 119187 (UCR); 2.IX.1977, 1♂ MEI 156351 (UCR); 6.IX.1977, 1♂ MEI 119163 (UCR), 30.IX.1977, Stone, M.W., Frommer, S., 1♂ MEI 119179 (UCR), San Timoteo Canyon [34.03, -117.204], 14.IX.1972, Wasbauer, M., Hardy, A., 3♀ MEI 072383–4, 073060 (CSCA), Malaise trap; Santa Rosa Plateau [33.529, -117.313], 686m, 23.V–1.VI.2000, Frommer, S.I., 1♀ MEI 139405 (UCR), chaparral; 7–19.IX.2000, Frommer, S.I., 1♀ MEI 152069 (UCR); Thousand Palms [33.82, -116.389], 31.III.1963, Parker, F.D., 1♂ MEI 170251 (UCDC); Wiley's Well Road at I-10 [33.606, - 114.908], 28.III.2001, Yanega, D., 1♂ UCR ENT 49590, 1♀ 49591 (UCR); Winchester [33.707, -117.084], 3.VI.1967, Icenogle, W., 1♂ MEI 119203 (MEIC); 10.VII.1967, 1♂ MEI 170246 (MEIC); 11.VIII.1967, 1♂ MEI 119162 (MEIC); 26.IV.1968, 1♀ MEI 119139 (MEIC); 20.V.1968, 1♂ MEI 119204 (MEIC); Lower Covington Flat [33.917, -115.917], 9.IV.1960, Knox, M., Sleeper, E., 1♂ MEI 112079 (CAS); Pinyon Wells [33.917, -115.917], 27.IV.1969, Sleeper, E.L., 1♂ MEI 112074 (CAS); Pleasant Valley [33.917, -115.917], 15.IV.1965, Sleeper, E.L., Jenkins, S.L., 1♀ MEI 112067 (CAS); 16.IV.1965, 1♂ MEI 112059 (CAS); 3.III.1967, 2♂ MEI 112022, 112046 (CAS); 8.IV.1967, 1♂ MEI 112023 (CAS). San Bernadino County. Clark Mountain, Clark Mountain [35.525, -115.588], 1.VII.1987, Nelson, G.H., 1♂ MEI 128579 (FSCA); 16.1 km NE Earp [34.267, -114.198], 17.IV.1964, Linsdale, D.D., 1♂ MEI 119161 (MEIC); Langston, R.D., 1♀ MEI 170269 (MEIC), blacklight trap; 17.7 km N Goffs, Lanfair Road [35.078, -115.059], 25.V.1977, Pinto, J.D., 1♂ MEI 119190 (UCR); 32.2 km W Trona [35.764, -117.662], 11.IV.1974, Morrison, R., Bezark, L.G., 1♂ MEI 119395 (UCR); E Highland [34.128, -117.211], 29.VIII.1947, Flock, R.A., 1♂ MEI 119214 (MEIC); Oak Glen Lodge [34.049, -116.947], 1524m, date?, collector?, 2♂ MEI 156360–1 (CAS); San Bernardino Mountains, Bear Valley, 2042m, Banks, N., 1♂ MEI 081095 (MCZ); Vidal Junction [34.188, - 114.575], 1.IV.1967, Blanc, F.L., 1♂ MEI 072651 (CSCA), at light. San Diego County. 1 km W Boulivard [32.652, -116.102], 1070m, 30.V–5.VI.2002, Irwin , M.E., Parker, F.D., 5♂ MEI 170227–31, 1♀ 170231 (MEIC), Malaise trap; 16.1 km S Julian, Inaja Indian Reservation [32.933, -116.603], 1204m, 11.IX.1969, Heppner, J.B., 1♂ MEI 119175, 1♀ 119239 (FSCA), blacklight; 19.3 km E Julian [33.132, -116.404], 12.IV.1990, Irwin , M.E., 1♂ MEI 119386 (MEIC); 2 km E Pine Valley [32.817, -116.517], 1300m, 1–5.VI.2002, Irwin , M.E., Parker, F.D., 1♀ MEI 152032 (MEIC), Malaise trap at seep in large wash; 6 km E Jacumba, near Desert View Tower [32.652, -116.102], 950m, 31.V–5.VI.2002, Irwin , M.E., Parker, F.D., 2♀ MEI 170234–5 (MEIC), Malaise trap; 6.4 km E Jacumba [32.617, -116.132], 15.IV.1964, Wilcox, J., 1♀ MEI 119072 (EMEC), 28.IV.1964, Wilcox, J., 2♂ MEI 119398, 119411 (EMEC); 21.IV.1966, Stone, M.W., 4♀ MEI 119229, 119409, 119069, 119397 (EMEC); 6.5 km NW Jacumba, near Bankhead Springs [32.652, - 116.102], 1070m, 30.V–5.VI.2002, Irwin , M.E., Parker, F.D., 3♀ MEI 170239, 170242–3 (MEIC), Malaise trap; 7.2 km E Borrego Springs [33.256, -116.311], 180m, 6.IV.1971, Fisher, E.M., 1♂ MEI 028338 (UCR); Borrego Springs [33.26, -116.38], 30.III.1960, Wasbauer, M., 1♂ MEI 170202 (UCDC); Coronado [32.654, - 117.158], 8.VII.1890, Blaisdell, F.E., 1♂ MEI 156359, 1♀ 080240 (CAS); Culp Canyon [33.208, -116.46], 1025m, 30.V–5.VI.2002, Irwin , M.E., Parker, F.D., 2♀ MEI 170236–7 (MEIC), Malaise trap; Descanso [32.835, -116.623], 1280m, date?, collector?, 2♂ MEI 170232–3 (MEIC), Malaise trap; El Cajon [32.795, - 116.962], 19.X.1983, Brown, J., 1♂ MEI 119171 (SDMC); Encinitas [33.05, -117.29], 4.VIII.1976, Huybensz, M., 1♀ MEI 119124 (MCZ); Jacumba [32.618, -116.189], 914m, 26.IV.1965, Wilcox, J., 1♂ MEI 119399, 1♀ MEI 119083 (EMEC); Laguna Mountains, Horsehaven Group Camp [32.897, -116.448], 1770m, 31.V.2002, Irwin , M.E., Parker, F.D., 1♀ MEI 170238 (MEIC), Malaise trap; Pine Valley [32.821, -116.528], 21.VI.1966, Sleeper, E.L., 1♀ MEI 112095 (CAS); San Diego, 5 km W Ranchita [33.213, -116.568], 1200m, 1–5.VI.2002, Irwin , M.E., Parker, F.D., 1♀ MEI 170241 (MEIC), Malaise trap; San Felipe Valley, Sissors Crossing [33.103, -116.487], 770m, 1–5.VI.2002, Irwin , M.E., Parker, F.D., 1♂ MEI 170203 (MEIC), Malaise trap; San Luis Rey [33.232, -117.323], 20.IV.1966, Wilcox, J., 1♂ MEI 119216 (EMEC). San Luis Obispo County. 32.2 km E Santa Maria [35.21, -120.148], 16.IV.1969, Pinto, Haines, 1♂ MEI 170200 (UCR). Santa Barbara County. 3.22 km E Solvang [34.596, -120.108], 28.VI.1965, Powell, J., 1♂ MEI 119217 (EMEC); San Rafael Mountains, Bluff Camp [34.674, -119.666], 29.VI.1959, Bohart, R.M., 1♂ MEI 119205 (UCDC). Ventura County. 1.6 km W Santa Susana [34.272, -118.722], 16.V.1964, Irwin , M.E., 1♂ MEI 119160 (MEIC), sandy wash. NEVADA. Humboldt County. Soldier Meadow [41.392, -119.163], 27.VII.1961, Parker, F.D., 1♂ MEI 119206 (MEIC), Lyon County. 3.2 km E Yerington [38.986, -119.133], 8.VIII.1973, Barclay, L.W., 1♂ MEI 076994 (NVDA), light trap; 4.8 km E Yerington [38.986, -119.119], 8.VIII.1973, Barclay, L.W., 2♂ MEI 076993, 076995 (NVDA), light trap. Nye County. 24.2 km W Tonopah [38.067, -117.447], 7.VII.1980, Hall, J.C., Hall, E.M., 1♂ MEI 078129 (UCR); Big Dune [36.647, -116.58], 29–30.IV.1982, Bezark, L.G., 2♂ MEI 119392, 119406 (BEZC), blacklight; Lava Dune, 30.IV.1982, Bechtel, R.C., Rust, R.W., 3♂ MEI 077013, 077017, 077019 (NVDA), blacklight; Mercury [36.661, -115.994], 24.VIII.1959, 1♂ MEI 113247 (USNM); 12.VII.1960, 1♂ MEI 113246 (USNM); 19.VI.1964, 1♂ MEI 113248 (USNM); Lockes [38.555, -115.774], 8.VII.1966, Allen, R.P., 1♂ MEI 072441 (CSCA). Pershing County. 11.3 km E Oreana [40.312, -118.188], 9.IX.1969, Haig, T.R., 1♂ MEI 072410 (CSCA). NEW MEXICO. 11.VIII.1966, Silberglied, R., 1♂ MEI 119207 (CUIC); Rosco, 8.IX.1958, Weems, H.V., Jr., 1♀ MEI 119137 (FSCA). Dona Ana County. Las Cruces [32.312, -106.778], 16.IV.1965, Medica, P.A., 1♂ MEI 051272 (NMSU); 30.V.1965, Medica, P.A., 1♀ MEI 051273 (NMSU). Hidalgo County. Animas [31.949, - 108.807], 27.V.1982, Peckham, D.J., 1♂ MEI 170266 (CUIC); Post Office Canyon, 9.VIII.1965, Ballmer, G.R., 1♂ MEI 119211 (MEIC). Lea County. [32.38, -103.722], 12.VII.1979, Delorme, D.R., McHugh, C.P., 1♂ MEI 076515 (TAMU), at light. Luna County. Deming [32.268, -107.759], 5.IX.1960, Marsh, P.M., 1♂ MEI 119212 (MEIC); Columbus [31.823, -107.641], 1237m, 29.VI.1997, Metz, M.A., 1♂ MEI 113891 (INHS), Malaise trap. Sandoval County. Frijoles Canyon [35.758, -106.308], 14.VI.1960, Burks, Kinzer, 1♂ MEI 113245 (USNM). TEXAS. Brewster County. 35.4 km S Marathon [29.887, -103.244], 8.IX.1960, Marsh, P.M., 1♂ MEI 119405, 1♀ 139398 (UCR).

Figures

FIGURES 56–63. Arenigena albiseta. 56. Tergite 8, epandrium, cerci. 57. Sternite 8, gonocoxite ventral view. 58. Gonocoxite, lateral view. 59. Gonostylus lateral view. 60. Aedeagus, dorsal view. 61. Aedeagus, ventral view. 62. Aedeagus, lateral view. 63. Female internal reproductive organs.

FIGURE 96–97. Distribution map. 96. Arenigena bajaensis, open circle, Ar. semitaria closed circle. 97. Arenigena marcida closed circle, Litolinga acuta open circle.