Arcyphysa diamontona, Wells & Contents, 2018

Wells, Alice & Contents, Arturs Neboiss Table Of, 2018, Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus, Zootaxa 4415 (1), pp. 1-44 : 27-28

Arcyphysa diamontona sp. nov.

( Figs 102–103, 106–107 , 116, 149, 151, 162–163 , 181–182)

Material examined. Holotype ♂, Victoria , Diamond Creek , 7 km SE of Gembrook, 31.i. 1979, Wells & Neboiss ( NMV [PT-1006]).

Paratypes. Victoria: 14♂ 10♀, data as for holotype ; 1♂ Noorinbee , 12.xi.1969, A. Neboiss ; 2♂ 2♀, 8 km NE Toolangi , 2.xii.1970, A. Neboiss ; 1♂ 1♀, Ada River , on Ada River road, 19.i.1979, NMV Survey Department ‘54’ ; 1♂ 2♀, LaTrobe River , 9.8 km W of Noojee, 21.i.1979, NMV Survey Department ‘55’ ; 2♂, Watts River , Fernshaw, 12 km NE of Healsville, 11.ii.1982; A. Neboiss & K. Walker ( NMV).

Other material, Victoria: 2♂, Gordon’s Bridge, Yea River, 6.i.1954, A. Neboiss; 1♂, Healsville, 7.ii.1954, A. Neboiss; 1♂, Noojee, 14.xi.1957, A. Neboiss; 1♂, Merrijig, Vic, 30.xi.1957, J. Landy; 2♂ 2♀, 7.xii.1957, J. Landy; 1♂, Merrijig, 15.i.1958, Edwards [PT-1095]; 1♂, Glenelg River, Chimney Pot Gap, Grampians, 16.xii.1966, A. Neboiss; 1♂, Glenelg River, Headwaters, 16.xii.1966, E.F. Riek ( ANIC, badly bleached specimen); 2♂ (? 38♀), Tooronga Falls, NE Noojee, 17.xii.1970, A. Neboiss; 4♂ 13♀, 3 km SE Taggerty, Little River, 6.i.1972, A. Neboiss; 1♂ 7♀, 3 km W Beenak, 7.i.1972, A. Neboiss; 31♀, Yea River, 7 km S of Glenburn, 1.xii.1972, A. Neboiss; 1♀, Aire River, 4 km? Beech Forest, 4.xii.1972, A. Neboiss & Malcolm; 1♀, Taggerty River, 4 km E of Marysville, 26.xii.1974, A. Neboiss; 2♀, Wingan River, 8 km S of Princes Hwy, 30.i.1975, A. Neboiss; 1♂, Jordan River at Jericho, 28.xi.1976, A.A. Calder; 2♂ 7♀, Matlock Creek off Thomson Portal Road, 14 km W of Aberfeldy, 10.ii.1977, A. Calder; 6♀, Tooronga River, 1 km S of Tooronga Rd, 28.xi.1978, NMV Survey Dept; 6♀, Tooronga Falls, 7 km NE of Noojee, 27.xi.1981, J. Morse & A. Neboiss; 1♂, Acheron River, 3.5 km ESE Narbethong, 14.i.1982, R. StClair; 2♂, Watts River, Fernshaw, 12 km NE of Healsville, 11.ii.1982, A. Neboiss & K. Walker; 8♀, O’Shannassy River, 20.i.1983, A. Neboiss; 2♂ 7♀, Vic, Matlock River off Thomson Portal Road, 14 km W of Aberfeldy, 10.ii.1977, A. Calder [TRI-39989].

Diagnosis. In general form, the male genitalia of this species are similar to those of Arc. angusta with strongly bowed gonopods, but with lateral extensions on fused abdominal tergites IX/X, in lateral view, broadly rounded and club-shaped, not narrowly lanceolate; a small down-curved spine is present medially on the apicolateral margins of abdominal segment IX; and on each gonopod, the harpago is acute apically, not rounded. The female terminal abdomen has apicolateral angles of sternite VIII narrowly digitiform.

Description. Length of each forewing ( Fig. 102 ): ♂ 9.0– 10.9 mm (n = 3), ♀ 10.3–11.8 mm (n = 7); forewing median cell length about 5x maximum width. Maxillary palps ( Fig. 103 ) with segments 2 and 3 about equal length, 1 and 4 shorter, 5 slightly longer than other four together.

Male ( Figs 106–107 , 162–163 ). Reticulate-walled internal sacs rounded, about 2/3 length of segments; lateral filaments short, about half length of segment. Genitalia: In lateral view with segment IX broadly concave apically; slender, spiny down-curved process on each lateral margin situated below level of phallus; tergites IX and X fused, in ventral view each side with elongate lateral extension of about equal width throughout length, on mesal margins slender process to about 2/3 length, in lateral view broadly rounded distally; gonopods in ventral view each with coxopodite stout, strongly curved, harpago short, narrow, in ventral view curved, tapered to acute apices directed mesad; phallic apparatus without parameres, stout basally, in lateral view tapered distally, down-turned.

Female. Abdomen ( Figs 181–182 ) terminating bluntly; pair of cerci and both pairs of terminal protuberances small; sternite VIII formed by 2 ventrolateral plates that cover inferior appendage receptacle grooves fully divided mesally, apicolateral angles extending dorsad in finger-like lobes, in lateral view rounded ventrally.

Distribution. Arcyphysa diamontona has been collected in areas immediately to the east and north-east of Melbourne, Victoria.

Figures

FIGURES 102–109. Arcyphysa gen. nov. spp., diagnostic features.102–103, Arc.diamontona sp. nov., Vic., Diamond Creek, 7 km SE of Gembrook [PT-1006], ♂: 102, right fore- and hind wings, dorsal, 103, maxillary palp. 104–109, ♂ genitalia. 104– 105, Arc. angusta (Banks 1939; NSW, Styx R. [PT-1005]): 104, ventral, and 105, left lateral. 106–107, Arc. diamontona sp. nov. [PT-1006]:106, ventral, and 107, left lateral.108–109, Arc.fraserensis sp. nov. (Qld, Fraser Island, Pile Valley, nr Central Station [PT-1125]):108, ventral,and 109, leftlateral.Scale bars: Fig.102 = 1 mm; Fig.103 = 0.5 mm.Abbreviations:ac= anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; fI, fII, fIII, fIV, fV = forks I–V; mc= median cell; har = harpago; ph = phallic apparatus; IX, X = abdominal segments IX, X

FIGURES 146–163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146–150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152–153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154–163, ♂ genitalia. 154–155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156–157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158–159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160– 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162–163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X.

FIGURES 179–187. Arcyphysa gen. nov. spp., diagnostic features. 179–182, ♀ terminal abdominal segments: 179–180, Arc. angusta (NSW, Styx R. [PT-1005]): 179, ventral, and 180, left lateral; 181–182, Arc. diamontona sp. nov. [PT-1006]: 181, ventral, and 182, left lateral. 183–187, ♂ reticulate-walled internal sacs of abdominal segments VI and VII: 183, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]), ventral; 184–185, Arc. acmea sp. nov. (Qld, Mt Bartle Frere,0.5 km Nof South Peak [PT-999]):184, rightlateral, and 185, ventral;186, Arc.anaplasis sp.nov. (Qld, 25 km along Mt Lewis Road, SW of Mossman [PT-1003]), ventral; 187, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]), ventral. Abbreviation:VIII = abdominal segment VIII.