Austropsyche Banks,

Wells, Alice & Contents, Arturs Neboiss Table Of, 2018, Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus, Zootaxa 4415 (1), pp. 1-44: 15

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Austropsyche Banks


Austropsyche Banks 

( Figs 49–101View FIGURES 49–58View FIGURES 59–69View FIGURES 70–83View FIGURES 84–101)

Austropsyche Banks 1939  , 496. Type species Austropsyche victoriana Banks 1939  , by monotypy.

Diagnosis. Austropsyche  is distinguished from Diplectrona  and Arcyphysa  gen. nov. mainly by the mesal processes on the coxopodites. As on the head of Diplectrona  , the anterolateral warts, if divided at all, are divided by only a poorly defined suture; in Austropsyche  the posterior portion is usually more or less fragmented. The forewings ( Figs 49, 50View FIGURES 49–58, 76–79View FIGURES 70–83) resemble those of Diplectrona  and Arcyphysa  in each having cell ac lanceolate and hind wings each with Sc and R1 curved towards the leading margin, but only slightly; the median cell length is equal to or about 2/3 that of the discoidal cell. In male genitalia, Austropsyche  , in common with Diplectrona  , may have sclerotised spines associated with the phallus; these are absent in Arcyphysa  .

Description. Head ( Figs 70–75View FIGURES 70–83) dorsally with 5 setal warts, anterolateral pair often incompletely divided, posterior part fragmented or at least irregular in appearance. Antennae with median suture on segments ( Fig. 80View FIGURES 70–83). Maxillary palpi ( Figs 51View FIGURES 49–58, 81View FIGURES 70–83) with segment 5 very long, equal to length of rest of palp, segment 3 about twice length of 4, but scarcely longer than 2, segments 2, 3 and 4 flattened, expanded and rounded distally ( Fig. 81View FIGURES 70–83). Forewing ( Figs 49, 50View FIGURES 49–58, 76–79View FIGURES 70–83) fork 4 equal to 5, A1+2+3 curving proximally so cell ac is lanceolate, not parallelsided, discoidal cell length equal to or about 2/3 length of median cell; hind wing median cell open, subcosta and radius running parallel to near middle of discoidal cell where they are connected by short cross-vein, and then separating, radius bending behind, but not so much as in Diplectrona  so as scarcely to approach discal cell and space not narrowed, costal area rather broader than in allied genera, not narrowed near middle, and with an oblique costal cross-vein before middle. Abdomen with lateral filaments ( Fig. 83View FIGURES 70–83) on segment V about length of segment or 0.75x length; paired reticulate-walled internal sacs ( Fig. 82View FIGURES 70–83) within abdominal segments VI and VII generally 1.5– 2x length of segments. Male genitalia characterised by mesal process on coxopodite of gonopods (e.g., Figs 52–53View FIGURES 49–58, 59–60, 62, 66, 68–69).

Remarks. In their unpublished MS redescribing Banks’ types, Ross and Morse (personal communication 1975) commented on the ‘unusual extensile slender apical portion’ of the phallus of Austropsyche victoriana  , on how in this respect Austropsyche  differs from any other Diplectroninae  genera. This structure is apparent in some of the museum specimens of Au. victoriana  , but not in all, and is not seen in any other species in the genus; possibly it depends on state at time of preservation, or method of preservation. On the basis of other character states, two distinct groups are recognised, referred to here as the Au. victoriana  Group, including Au. victoriana  , Au. bifurcata (Kimmins, in Mosely & Kimmins 1953)  , and Au. kaputar  sp. nov.; and the Au. acuta  Group, comprising Au. acuta  sp. nov. and Au. ambigua  sp. nov. Placement of a 6th species, Au. morana  sp. nov., is enigmatic; it appears to combine some male genitalic features of both groups, but in other features is highly autapomorphic.

Distribution. Found from the Grampians of south-western Victoria to south-eastern Queensland; no species of Austropsyche  is known to occur in Tasmania.












Austropsyche Banks

Wells, Alice & Contents, Arturs Neboiss Table Of 2018


Banks 1939

Austropsyche victoriana Banks 1939

Banks. Diagnoses 1939