Arcyphysa carnarvona, Wells & Contents, 2018

Wells, Alice & Contents, Arturs Neboiss Table Of, 2018, Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus, Zootaxa 4415 (1), pp. 1-44 : 33-34

Arcyphysa carnarvona sp. nov.

( Figs 126–127 , 147 , 171–173)

Material examined. Holotype ♂, Queensland, Carnarvon Gorge National Park , 25°15'S 148°24'E, 12.xi.1990, G. Theischinger ( NMV [TRI-39506]).

Paratypes, Queensland: 2♂ 3♀, collected with holotype [TRI-39506].

Diagnosis. This species groups with Arc. crescentina, Arc. sybillae, Arc. caldera , and Arc. hugginsi , having, in the males, parameres on the phallus. Among these species, males most closely resemble those of Arc. hugginsi and Arc. caldera in having two pairs of long apicolateral processes on segment IX. Arcyphysa carnarvona most closely resembles Arc. hugginsi in having the outermost pair heavily sclerotised, but in Arc. carnarvona the processes are sinuous, crossed medially, crenate, and slightly truncate apically, while in Arc. hugginsi the processes curve gently, swell distally to be bulbous and then converge without crossing toward pointed, beak-like apices.

Description. Length of each forewing: ♂ 8.7 mm (n = 3), ♀ 9.6 mm (n = 3).

Male. In abdomen, glandular sacs ovoid, about one segment long; filaments on segment V short, about 1/4 length of segment. Genitalia ( Figs 126–127 , 171–173 ): Abdominal segment IX apicolaterally with 2 processes on each side, outermost heavily sclerotised, sinuous, apically crenate, and innermost, more dorsal process pale and terminating in dense spines; tergite IX slightly sclerotised, evenly rounded distally, fused with membranous segment X; segment X in dorsal view rounded distally, shallowly cleft medially; gonopods each with coxopodite in ventral view about 5x maximum width, harpago short, curved inward, apically abruptly reduced to acute spur, in lateral view bifid apically; phallic apparatus expanded basally with very short, conical, apically sclerotised paramere on each side, shorter than in D. sybillae .

Female. Terminalia almost 'lotus-like' in ventral view, with sternite VIII in form of 2 well-separated valves, slightly divided distally and laterally produced in narrow distal lobes; sternite IX with paired leaf-like sclerotisations laterally; segment X with cerci and paired apical papillae well developed.

Distribution. Known only from Carnarvon Gorge in the Brigalow country of southern Queensland.

Figures

FIGURES 117–131. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 117–118, Arc. caldera sp. nov. (Qld, The Crater,Atherton [PT-1040]):117, ventral; 118, leftlateral.119–120, Arc.crescentinasp.nov. (Qld, TullyFalls [PT-1002]):119, ventral; 120, left lateral. 121–122, Arc.hugginsi sp. nov. (Qld, Bellenden Ker Range [PT-1000]): 121, ventral; 122, left lateral. 123–125, Arc. sybillae sp. nov. (Qld, Bloomfield River [PT-1016]): 123, ventral; 124, left lateral; 125, dorsal. 126–127, Arc. carnarvona sp. nov. (holotype; Qld, Carnarvon Gorge National Park): 126, ventral; 127, left lateral. 128–129, Arc. naumanni sp. nov. (holotype; Qld, 11 km W by N Bald Hill, McIllraith Range): 128, ventral; 129, left lateral. 130–131, Arc.lovedayi sp. nov. (holotype): 130, ventral; 131, left lateral.

FIGURES 146–163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146–150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152–153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154–163, ♂ genitalia. 154–155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156–157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158–159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160– 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162–163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X.

FIGURES 164–178. Arcyphysa gen. nov. spp., ♂ genitalia diagnostic features. 164–165, Arc. lovedayi sp. nov. (holotype): 164, ventral; 165, left lateral. 166–167, Arc. crescentina sp. nov. (Qld, Tully Falls [PT-1002]): 166, ventral; 167, left lateral. 168–169, Arc. hugginsi sp. nov. (Qld, Bellenden Ker Range [PT-1000]): 168, ventral; 169, left lateral. 170, Arc. sybillae sp. nov. (Qld, Bloomfield River [PT-1016]), ventral. 171–173, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge): 171, ventral; 172, left lateral;173, dorsal. 174–175, Arc.anaplasis sp. nov. (Qld, 25 km along Mt LewisRoad, SW of Mossman [PT-1003]): 174, ventral; 175, left lateral.176, Arc.acmea sp. nov. (Qld, Mt Bartle Frere, 0.5 km N of South Peak [PT-999]), ventral.177– 178, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]): 177, ventral; 178, left lateral.