Scydmaenichnus Reitter
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publication ID |
https://doi.org/ 10.11646/zootaxa.3630.1.2 |
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publication LSID |
lsid:zoobank.org:pub:A338EE37-E8AC-4AC3-8D6B-D0177FB680BF |
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DOI |
https://doi.org/10.5281/zenodo.5696216 |
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persistent identifier |
https://treatment.plazi.org/id/03D687E5-B05C-9A75-BDD1-43A7118AFF66 |
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Plazi |
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scientific name |
Scydmaenichnus Reitter |
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Subgenus Scydmaenichnus Reitter
Scydmaenichnus Reitter, 1905: 99 (as subgenus of Stenichnus ). Type species: Stenichnus fossifrons Reitter, 1905 (designated by Franz in Newton & Franz (1998)).
Lectotype designation. Redescription of this subgenus given below is based on the morphology of a single male type specimen of Stenichnus fossifrons Reitter , the type species of Scydmaenichnus , preserved at HNHM. This specimen ( Figs. 45–51 View FIGURES 45 – 51 ), having a status of a syntype (Reitter (1905) mentioned more than one specimen in his original description of S. fossifrons ), is here designated lectotype, in order to ensure the stability of nomenclature and provide a unique name-bearing type for Stenichnus fossifrons .
Revised diagnosis. A subgenus of Stenichnus showing the following differences in relation to Stenichnus s. str.: head in males modified, with buckling frons (non-modified in Stenichnus s. str.); setose impressions of mesoventrite broadly separated from mesocoxal cavities (narrowly separated in Stenichnus s. str.); each elytron with two rudimentary foveae (one deep and one rudimentary in Stenichnus s. str.); and hind wings absent (present in Stenichnus s. str.).
Redescription. Body ( Fig. 45 View FIGURES 45 – 51 ) strongly convex, elongate and relatively slender, with long appendages, vestiture moderately long and dense.
Head ( Figs. 46–48 View FIGURES 45 – 51 ) typical for Stenichnus , with short tempora, eyes located posteriorly, fronto-clypeal groove present; trapezoidal submentum with long posterior margin laterally demarcated from hypostomae; hypostomal ridges complete. In males frons between eyes and antennal insertions expanded anteriorly and forming large protuberance demarcated from vertex by impression ( Fig. 47 View FIGURES 45 – 51 ; fr). Antennae ( Fig. 45 View FIGURES 45 – 51 ) slender, gradually thickening towards apices.
Prothorax ( Figs. 45, 48 View FIGURES 45 – 51 ) in dorsal view with rounded anterior and lateral margins of pronotum, with transverse row of four small ante-basal foveae, without lateral marginal carinae or edges; hypomera ( Fig. 48 View FIGURES 45 – 51 ; hy) not demarcated from notum. Prosternum ( Fig. 48 View FIGURES 45 – 51 ) with short basisternal part ( Fig. 48 View FIGURES 45 – 51 ; bs), procoxal cavities ( Fig. 48 View FIGURES 45 – 51 ; pcc) with anterior marginal carina and separated in middle by fine prosternal intercoxal carina; notosternal sutures ( Fig. 48 View FIGURES 45 – 51 ; nss) complete.
Mesoventrite ( Fig. 48 View FIGURES 45 – 51 ;) with narrow anterior ridge, lateral asetose impressions ( Fig. 48 View FIGURES 45 – 51 ; ai) separated in middle and lateral setose impressions ( Fig. 48 View FIGURES 45 – 51 ; si) narrowly separated in middle and broadly separated from mesocoxal cavities ( Fig. 48 View FIGURES 45 – 51 ; mscc); mesocoxal projection ( Fig. 48 View FIGURES 45 – 51 ; mcp) with carina and with posterior lobe ( Fig. 48 View FIGURES 45 – 51 ; pl) bearing dense setae along posterior margin; mesoventral intercoxal process ( Fig. 48 View FIGURES 45 – 51 ; msvp) narrow, not fused with metaventrite.
Metaventrite ( Fig. 48 View FIGURES 45 – 51 ) broader than long, with metaventral anterior process ( Fig. 48 View FIGURES 45 – 51 ; mtap) concealed by dense setae; metaventral intercoxal process ( Fig. 48 View FIGURES 45 – 51 ; mtvp) broad and short, with distinct median notch.
Elytra ( Fig. 45 View FIGURES 45 – 51 ) oval, with highly reduced humeral calli, narrow at base, each elytron with two rudimentary basal foveae barely noticeable in dry-mounted specimens. Hind wings absent.
Legs ( Fig. 45 View FIGURES 45 – 51 ) long; femora clavate, tibiae slender, tarsi strongly elongate.
Abdominal sternites VII and VIII separated by indistinct suture.
Aedeagus ( Figs. 49–50 View FIGURES 45 – 51 ) with structures typical for Stenichnus , ie., with ventral projection ( Fig. 49 View FIGURES 45 – 51 ; vp), mesoventral projection ( Fig. 49 View FIGURES 45 – 51 ; mvp) and lightly sclerotized endophallus ( Fig. 49 View FIGURES 45 – 51 ; end) with denticulate surface; parameres free and slender, with apical setae.
Distribution and composition. Five species of Stenichnus (Scydmaenichnus) are distributed in Cyprus, Israel and Turkey. Some old records refer to " Syria " (Davies 2004), a territory at that time different from the present country, so these data may concern adjacent countries of the Middle East.
Remarks. The large number of characters shared with Stenichnus s. str. and only a few minor differences found during the present study ( Table 1 View TABLE 1 ) justify the placement of Scydmaenichnus as a subgenus. In the available material (single and fragile type specimen) it was not possible to examine the structure of mandibles and prementum. The mandibles seem to be falciform in S. fossifrons , but their proximal parts were not visible in a temporary transparent mount. The broadly separated bases of maxillary palps suggest that the prementum is modified in a similar way as in Stenichnus s. str., but further morphological study is necessary to analyze all important structural details of the mouthparts.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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