Plusioglyphiulus foveatus, Golovatch & Geoffroy & Mauriès & Den Spiegel, 2009

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Den Spiegel, Didier Van, 2009, Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae), Zoosystema 31 (1), pp. 71-116 : 83-86

Plusioglyphiulus foveatus n. sp.

( Figs 8-10)

TYPE MATERIAL. — Laos. Luang Prabang Prov., 19.88°N, 102.14°E, W of Tham , 3.VIII.1992, leg. J.- P. Besson, holotype ♂ ( MNHN GA056) ; 1 ♀ paratype ( MNHN GA056); 1 ♂ paratype ( SEM).

ETYMOLOGY. — To emphasize the foveate apex of the posterior gonopods.

DIAGNOSIS. — Differs from congeners by certain details of gonopod structure (characteristic shapes of the different outgrowths and processes), coupled with the mixostichic carinotaxy of the collum, the isostichic carinotaxy of other body segments, the convex paraprocts, the broadly rounded epiproct and the presence of spots dorsad of the ocellaria.

DESCRIPTION

Length 28-35 mm, width 1.3-1.9 mm, collum and midbody segments being equally broadest.

Coloration whitish to yellow-brown; collum and, partly, head and segment 2 entirely pale; tips of most of tergal crests/tubercles, as well as a narrow axial line and a slightly broader lateral stripe level to ozopores pale grey-brown to brown; ocellaria dark brown with pale to dark brown, widely separated spots dorsally.

Body with 49-66p+4-1a+T. Length of holotype about 30 mm, width 1.3 mm, with 58p+2a+T.

Antennae rather short and clavate ( Fig. 10A), antennomeres 6 and 7 each with a considerable group of bacilliform sensilla distodorsally; similar but minute sensilla present also on antennomere 7 ( Fig. 9C). Ocellaria black, ovoid in shape, with 10-13 ocelli in 5-7 longitudinal rows. Gnathochilarium oligotrichous.

Postcollar constriction evident, but collum and segment 2 relatively modestly enlarged ( Fig. 8B). Carinotaxy formula of collum:ta/t+2p/t+3p/t+4p/t/ t+ta/5/t+(ta)/t+VI(p)/t+pp/t/t+ma/t ( Fig. 8A, B). Carinotaxy of metaterga 2-4, 7/7+m/m+7/7, all crests being longitudinal; formula of metaterga 5 to 15, 3/3+I/i+3/3/3+m/m+3/3/3+I/i+3/3; formula of subsequent metaterga usually 3/3+I/i+3/3/3+m/ (m)/m+3/3/3+I/i+3/3; that of apodous segments, 3+4/4+m/m+4/4+3 ( Fig. 8); all crests and tubercles rather high, poriferous cones (I) much higher, nearly as high as broad, but median ones (m) very low ( Fig. 9B). Dorsal crests on several posteriormost segments considerably higher than others ( Fig. 8E). Midbody segments circular in cross-section ( Fig. 9B). Pleural regions of segments 2-4 conspicuously expanded, flap-shaped, especially so on segment 3 ( Fig. 8A). An evident, transverse pleural ridge behind gonopod aperture on male segment 7, with rounded flaps bent abruptly caudad. Tegument and limbus as in P. grandicollis or P. ampullifer n. sp. Epiproct broadly rounded and cariniform apically, with 2+2 paramedian tubercles at midway; paraprocts regularly convex, not flattened; hypoproct emarginate at caudal margin ( Fig. 9A).

Legs short, on midbody segments about ⅔ as long as segment diameter ( Fig. 9B). Claw at base with a strong accessory spine almost half as long as claw itself ( Figs 9D; 10E). Tarsi, tarsal setae and male legs 1-3 ( Fig. 10 B-D) as in P. grandicollis .

Anterior gonopods ( Fig. 10F) much like in P. ampullifer n. sp., but processes cxp1 large, non-contiguous, flattened sagittally and each carrying a short medial uncus apically; processes cxp2 slightly curved, slender and bifid; telopodites (te) simple, movable, elongate-subconical, setose laterally in distal half. Posterior gonopods ( Fig. 10G) relatively simple, with a pair of strongly reduced telopodites (te) laterally at base; coxite with short denticles (d) instead of arms, fovea-shaped apically.