Enema pan ( Fabricius, 1775 )

Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 199

Enema pan ( Fabricius, 1775)

Scarabaeus pan Fabricius 1775: 5 ( India)

= Scarabaeus chorinaeus Fabricius 1775: 5 ( Brazil)

= Enema infundibulum Burmeister 1847: 234 ( Brazil - LECTOTYPE)

Distribution. Guyana: Erichson 1848: 561 (as E. infundibulum ). French Guiana: Endrödi 1976: 32, 61 (as E. pan ; as Scarabaeus chorinaeus (treated as a synonym of Megaceras lycaon (now M. philoctetes ))), 1985a: 579, 580; Dechambre 1979c: 168; Lachaume 1992: 46; Brûlé et al. 2011a: 191; Duranton 2011: 22; Gasca-Álvarez and Ratcliffe 2011: 25; Ponchel 2011: 61. “Guianas”: Burmeister 1847: 234 (as E. infundibulum ); Blackwelder 1944: 256 (as E. infundibulum (treated as a synonym of E. pan var. enema )). Brazil (NA): Fabricius 1775: 5 (as Scarabaeus chorinaeus ); Burmeister 1847: 234 (as E. infundibulum ); Arrow 1937b: 65; Blackwelder 1944: 256; Endrödi 1973: 15 (designation LECTOTYPE Enema infundibulum ), 1976: 32, 61, 1985a: 579, 580; Dechambre 1979c: 168; Lachaume 1992: 46; Gasca-Álvarez et al. 2008: 13 (NA); Gasca-Álvarez and Fonseca 2009: 721 (NA); Gasca-Álvarez and Ratcliffe 2011: 25. Other: Fabricius 1775: 5 ( India - as Scarabaeus pan ); Arrow 1937b: 65 ( Colombia, Costa Rica, Mexico, Nicaragua, Paraguay, Peru); Blackwelder 1944: 256 ( Argentina, Colombia, Costa Rica, Mexico, Nicaragua, Paraguay, Peru); Endrödi 1976: 32, 61 ( Bolivia, Colombia, Ecuador, Paraguay, Peru - as E. pan ; Peru - as Scarabaeus chorinaeus (treated as a synonym of Megaceras lycaon (now M. philoctetes ))), 1985a: 579, 580 ( Bolivia, Colombia, Ecuador, Paraguay, Peru); Dechambre 1979c: 168 ( Bolivia, Colombia, Ecuador, Paraguay, Peru); Lachaume 1992: 46 ( Bolivia, Colombia, Ecuador, Paraguay, Peru); Gasca-Álvarez et al. 2008: 13 (Mexico–Paraguay); Gasca-Álvarez and Ratcliffe 2011: 25 ( Bolivia, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru); Ratcliffe et al. 2013: 369 (Mexico– South America); Dupuis 2016b: 108 (see E. endymion ).

Note. According to Dupuis (2016b: 108), Enema pan does not occur in French Guiana, with all records of this species in French Guiana being based on misidentifications of E. endymion . We doubt if E. pan is present in the Guianas, especially since we are also unaware of any records from Venezuela. It would be interesting to see if the old record from Guyana in Erichson (1848: 561) and the specimens identified as E. pan found in Brazil north of the Amazon River actually belong to this species, or if they are also misidentified E. endymion . Because of the current identification of the specimens from the Brazilian part of the research area, we regard E. pan as potentially occurring in the Guianas.