Cryptachaea pilar Santanna, 2019

Santanna, Manoela, Rodrigues, Everton Nei Lopes, Cizauskas, Igor & Brescovit, Antonio Domingos, 2019, On the spider genus Cryptachaea from Peru, Bolivia and cave environments in Brazil: a new species, additional descriptions and new records (Araneae, Theridiidae), Zootaxa 4646 (2), pp. 271-292: 273-276

publication ID

https://doi.org/10.11646/zootaxa.4646.2.4

publication LSID

lsid:zoobank.org:pub:E031D297-36C5-46D3-A7B0-7355719AAD64

persistent identifier

http://treatment.plazi.org/id/03D80F63-FFB6-FFA1-FF0F-7BE1FBC1FE2D

treatment provided by

Plazi

scientific name

Cryptachaea pilar Santanna
status

 

Cryptachaea pilar Santanna   & Rodrigues, new species

Figs. 1–4 View FIGURES 1–4 , 9, 10 View FIGURES 9–14 , 20–25 View FIGURES 20–25 , 32 View FIGURE 32

Types. Male Holotype, BRAZIL: Minas Gerais, Morro do Pilar, Cave CAV-0018 (19°10’13’’S; 43°23’46’’W), 13– 17.II.2012, L. Tunes coll., manual collection ( IBSP 189596 View Materials ) GoogleMaps   . Paratypes. BRAZIL: Same data as holotype, 1F ( IBSP 189596 View Materials )   ; Same data as holotype, except 03–14/VI/2013, Equipe Carste coll., 1M, 2F ( IBSP 189609 View Materials )   ; Minas Gerais, Sete Lagoas, Cave ILCOM-14 (19°29’47”S; 44°13’16”W), 05–14.VIII.2013, Equipe Carste coll., manual collec- tion, 1F ( IBSP 189573 View Materials ) GoogleMaps   ; Minas Gerais, Santa Maria de Itabira, Cave Morro Escuro (19°24’14”S; 43°08’36”W), 24.VI–05.VII.2013, Equipe Carste coll., manual collection, 1F ( IBSP 189621 View Materials ) GoogleMaps   .

Etymology. The species name is a noun in apposition taken from the type locality.

Diagnosis. The male palpus of Cryptachaea pilar   new species differs from the other species of the genus by their very large, subquadrangular tegulum that ventrally covers all the embolus ( Figs. 1 View FIGURES 1–4 , 20, 22 View FIGURES 20–25 ), and apex of the cymbium without projections ( Figs. 1, 2 View FIGURES 1–4 , 20–22 View FIGURES 20–25 ). The female have two circular copulatory openings ( Figs. 3, 4 View FIGURES 1–4 , 25 View FIGURES 20–25 ), as Cryptachaea cidae   Rodrigues & Poeta, 2015 (see Rodrigues & Poeta, 2015: fig. 1b), but differ from this species by the oval spermathecae ( Figs. 3, 4 View FIGURES 1–4 ), more separated from each other and fertilization ducts inserted submedianally in the spermathecae ( Fig. 4 View FIGURES 1–4 ), posteriorly in C. cidae   .

Description. Male holotype ( IBSP 189596): Total length 1.57. Carapace: length 1.57, width 0.71. Clypeus height: 0.12. Sternum: length 0.37, width 0.39. Chelicerae: length 0.31, width 0.10. Abdomen: length: 0.98, width 0.84, height 1.00. Leg formula I/II/III/IV. Segment length (I/II/III/IV): femora 1.11/0.77/0.65/0.81; patellae+tibiae 1.13/0.81/0.58/0.84; metatarsi+tarsi 1.34/1.00/0.73/1.32; total: 3.59/2.60/1.97/1.65. Palp: femora 0.21, patellae 0.10, tibiae 0.06, cymbium 0.18. All eyes are the same size. Eyes with dark borders ( Fig. 9 View FIGURES 9–14 ). Carapace oval, yellow with dark margins ( Fig. 9 View FIGURES 9–14 ). Clypeus yellow, laterally concave. Chelicerae ( Fig. 24 View FIGURES 20–25 ) yellow. Labium yellow, subquadrangular. Endites yellow, subquadrangular. Ocular area yellow, slightly darker between median eyes ( Fig. 9 View FIGURES 9–14 ). Sternum yellow, with dark dots. Legs yellow, except distal femur apex with dark brown spot. Presence of a comb of bristles distributed serially on the femur I, in ventral view. Abdomen oval ( Fig. 9 View FIGURES 9–14 ), dorsally grayish, with median and posterior dark brown spots; ventrally yellow, with dark brown dots. Spinnerets brown. Palp with lamellar conductor ( Figs. 1 View FIGURES 1–4 , 20, 21 View FIGURES 20–25 ). Embolus with two apical processes, one of them more robust ( Figs. 2 View FIGURES 1–4 , 21 View FIGURES 20–25 ), the minor process serves as a hook, with prolateral coupling, in the concavity of the alveolus of the cymbium ( Fig. 2 View FIGURES 1–4 ). Tegulum subquadrangular, large and robust ( Figs. 1 View FIGURES 1–4 , 20, 22 View FIGURES 20–25 ). Cymbium oval ( Fig. 2 View FIGURES 1–4 ), with tarsal organ circular and large ( Figs 21, 23 View FIGURES 20–25 ), and with stridulatory ridges in the apex of cymbium ( Fig. 23 View FIGURES 20–25 ).

Female paratype ( IBSP 189573). Total length 2.07. Carapace: length 0.98, width 0.92. Clypeus height: 0.14. Sternum: length 0.54, width 0.52. Chelicerae: length 0.31, width 0.12. Abdomen: length: 1.26, width 1.09, height 1.09. Leg formula I/IV/II/III. Segment length (I/II/III/IV): femora 1.21/0.96/0.65/1.19; patellae+tibiae 1.42/1.02/0.77/1.26; metatarsi+tarsi 1.68/0.86/0.67/1.34; total: 4.32/2.85/2.1/3.80. Palp: femora 0.21, patellae 0.16, tibiae 0.12, tarsi 0.14. PME larger than the other eyes; other eyes are the same size. Color of carapace ( Fig. 10 View FIGURES 9–14 ), legs and abdomen as in male. Epigynum with epigynal plate wider than long ( Figs. 3 View FIGURES 1–4 , 25 View FIGURES 20–25 ), with two spherical copulatory openings ( Figs. 3 View FIGURES 1–4 , 25 View FIGURES 20–25 ), two oval spermathecae ( Fig. 3 View FIGURES 1–4 ), and short fertilization and copulatory ducts ( Fig. 4 View FIGURES 1–4 ).

Variation. Males (n=2): Total length 1.57–1.89. Carapace: length 0.71–0.73, width 0.67–0.73. Femur leg I 1.11–1.26. Females (n= 5): Total length 2.07–2.73. Carapace: length 0.98–1.36, width 0.81–1.19. Femur leg I 1.21– 1.57.

Additional material examined: BRAZIL. São Paulo, Altinópolis, Sertãozinho de Baixo, Cave Gruta do Paredão (21°3'48''S; 47°25'47''W), 4 F, P. Gnaspini et al. coll., 21.XI.1998 ( IBSP 23841 View Materials ) GoogleMaps   ; Minas Gerais, Cordisburgo, Cave Gruta do Salitre (19°7'18''S; 44°21'5''W), 1 F, P. Gnaspini coll., 11.IX.2001 ( IBSP 71838 View Materials ) GoogleMaps   ; Cave Gruta da Morena (19°10'9''S; 44°19'54''W), 4 F, P. Gnaspini coll., 12.IX.2001 ( IBSP 71834 View Materials ), 8F, 11–12.X/2007 ( IBSP 132138 View Materials ), 3F ( IBSP 132145 View Materials ), 1F ( IBSP 132149 View Materials ), 5F ( IBSP 132150 View Materials ), 3F ( IBSP 132164 View Materials ), 5F ( IBSP 132165 View Materials ) all collected by Equipe Disciplina Cavernas 2007 GoogleMaps   ; Mariana: Cave Furna Dois   (20°13'35''S; 43°11'29''W), 4 F, F. Tulio coll., IX.2002 ( IBSP 39738 View Materials ), 2F, 14.VII.2002 ( IBSP 39778 View Materials ) GoogleMaps   ; Mina Conta História, Cave Caverna CH-033 1M, 23–30.IX.2008 ( IBSP 146072 View Materials ), 2F, 28–30.IV.2009 ( IBSP 146070 View Materials )   ; Cave Caverna CH-035, 1F, 28–30.IV.2009 ( IBSP 146079 View Materials ), all collected by F. Pellegatti et al.   ; Fábrica Nova, Cave Caverna FN-001 (20°13'41''S; 43°25'50''W), 2F, 05–09.V.2008 ( IBSP 199865 View Materials ), 3F ( IBSP 146107 View Materials ), 3F, 05–09.V.2009 ( IBSP 146136 View Materials ), 4F ( IBSP 146145 View Materials ), 3M, 3F ( IBSP 146146 View Materials ) GoogleMaps   ; Cave Caverna FN-002 (20°13'39''S; 43°25'53''W), 2F, 16–21.I.2009 ( IBSP 146116 View Materials ), 3F ( IBSP 146120 View Materials ) GoogleMaps   ; Cave Caverna FN-005 (20°13'19''S; 43°26'4''W), 2F, 05–09.V.2009 ( IBSP 146130 View Materials ) GoogleMaps   ; Cave Caverna FN- 006 (20°13'8''S; 43°25'51''W), 1F, 05–09.V.2009 ( IBSP 146148 View Materials ), 1F ( IBSP 146155 View Materials ), 1M, 2F ( IBSP 146158 View Materials ), 1F ( IBSP 146159 View Materials ), 1F ( IBSP 146167 View Materials ), 2M ( IBSP 146172 View Materials ), 1F ( IBSP 146175 View Materials ), 2M, 6F ( IBSP 146178 View Materials ) GoogleMaps   ; Cave Caverna FN-009 (20°12'28''S; 43°26'20''W), 1M, 16–21.I.2009 ( IBSP 146187 View Materials ) GoogleMaps   ; Cave Caverna FN-010 (20°12'29''S; 43°26'20''W), 2F, 05–09.V.2009 ( IBSP 146196 View Materials ) GoogleMaps   ; Cave Caverna FN-012 (20°12'30''S; 43°26'20''W), 1F, 16–21.I.2009 ( IBSP 146208 View Materials ), 2F ( IBSP 146213 View Materials ) GoogleMaps   ; Cave Caverna FN-013 (20°12'28''S; 43°26'19''W), 1M, 1F, 05–09.V.2009 ( IBSP 146223 View Materials ), 3F ( IBSP 146229 View Materials ), 1M, 3F ( IBSP 146238 View Materials ), 1M, 5F ( IBSP 146240 View Materials ) GoogleMaps   ; Cave Caverna FN-015 (20°12'22''S; 43°26'10''W), 2F, 05–09.V.2009 ( IBSP 146257 View Materials ), all collected by R. Andrade et al. GoogleMaps   ; Cave Caverna C-4 (20°11'25''S; 43°30'34''W), 1F, Bessi et al. coll., 16–17.IV.2009 ( IBSP 149757 View Materials ) GoogleMaps   ; Cave Caverna C-2 (20°11'24342''S; 43°30'34''W), 1F, Bessi et al. coll., 04–05.VIII.2009 ( IBSP 149797 View Materials )   ; Ouro Preto, Mina Conta História, Cave Caverna CH-007 (20°14'36726''S; 43°31'3''W), 1F, 23–30.IX.2008 ( IBSP 145932 View Materials )   ; Cave Caverna CH-010 (20°14'41496''S; 43°31'1''W), 1F, 23–30.IX.2008 ( IBSP 145896 View Materials )   ; Cave Caverna CH-011 (20°14'9''S; 43°30'39''W), 1F, 28–30.IV.2009 ( IBSP 145946 View Materials ), 1F ( IBSP 145949 View Materials ) GoogleMaps   ; Cave Caverna CH-013 (20°14'11''S; 43°30'34''W), 1F, 28–30.IV.2009 ( IBSP 145954 View Materials ) GoogleMaps   ; Cave Caverna CH-029 (20°13'54''S; 43°30'46''W), 1F, 23–30.IX.2008 ( IBSP 146047 View Materials ) GoogleMaps   ; Cave Caverna CH- 031 (20°14'2''S; 43°30'27''W), 1F, 23–30.IX.2008 ( IBSP 146063 View Materials ) GoogleMaps   ; Barão de Cocais, Cave Gruta Córrego Vieira 1 F, R. Andrade coll., 02–24.XI.2007 ( IBSP 127607 View Materials ), 1F ( IBSP 127609 View Materials )   ; Caeté: Cave Caverna AP_047 (20°2'12''S; 43°41'32''W), 1F, 12–21.XI.2009 ( IBSP 179057 View Materials ), 2M, 2F ( IBSP 179058 View Materials ), 1M, 1F ( IBSP 179059 View Materials ), 2F, 13–17.IV.2010 ( IBSP 179078), 1F ( IBSP 179079), 1F ( IBSP 179080) GoogleMaps   ; Cave Caverna AP_049 (20°2'9672''S; 43°41'30''W), 1F, 13–17.IV.2010 ( IBSP 179082)   ; Cave Caverna AP_050 (20°1'54732''S; 43°41'37''W), 5F, 12–21.XI.2009 ( IBSP 179054 View Materials ), 1F ( IBSP 179055 View Materials ), 1M, 1F, 13–17.IV.2010 ( IBSP 179083 View Materials )   ; Cave Caverna AP_054 (20°1'41574''S; 43°40'53''W), 1F, 12–21.XI.2009 ( IBSP 179052 View Materials ), 2F, 13–17.IV.2010 ( IBSP 179094 View Materials )   ; Cave Caverna AP_057 (20°1'36''S; 43°41'9''W), 1F, 12–21.XI.2009 ( IBSP 179076 View Materials ), 2F ( IBSP 179077 View Materials ), 1F, 13–17.IV.2010 ( IBSP 179084 View Materials ) GoogleMaps   ; Cave Caverna AP_058 (20°1'35''S; 43°41'9''W), 1M, 1F, 12–21.XI.2009 ( IBSP 179071 View Materials ), 1F ( IBSP 179072 View Materials ), 1F ( IBSP 179073 View Materials ), 1F 13–17.IV.2010 ( IBSP 179086 View Materials ), 1F ( IBSP 179085 View Materials ) GoogleMaps   ; Cave Caverna AP_059 (20°1'35''S; 43°41'9''W), 1F, 12–21.XI.2009 ( IBSP 179060 View Materials ) 1M, 1F ( IBSP 179061 View Materials ), 2F ( IBSP 179062 View Materials ), 1F ( IBSP 179063 View Materials ), 2F, 13–17.IV.2010 ( IBSP 179087 View Materials ),1F ( IBSP 179088 View Materials ), 1F ( IBSP 179089 View Materials ), 1F ( IBSP 179090 View Materials ) GoogleMaps   ; Cave Caverna AP_058 (20°1'35''S; 43°41'9''W), 1F, 13–17.IV.2010 ( IBSP 179081 View Materials ) GoogleMaps   ; Cave Caverna AP_061 (20°1'32''S; 43°41'8''W), 1F, 12–21.XI.2009 ( IBSP 179070 View Materials ), 1F, 13–17.IV.2010 ( IBSP 179091 View Materials ), 1F ( IBSP 179092 View Materials ) GoogleMaps   ; Cave Caverna AP_063 (20°1'31''S; 43°41'7''W), 1F, 13–17.IV.2010 ( IBSP 179093 View Materials ) GoogleMaps   ; Cave Caverna AP_065 (20°1'24''S; 43°41'4''W), 1F, 12–21.XI.2009 ( IBSP 179074 View Materials ), 1F ( IBSP 179075 View Materials ) GoogleMaps   ; Cave Caverna AP_064 (20°1'29784''S; 43°41'7''W), 1F, 12– 21.XI.2009 ( IBSP 179069 View Materials )   ; Cave Caverna AP_069 (20°0'36''S; 43°40'10''W), 2F, 12–21.XI.2009 ( IBSP 179067 View Materials ), 1M ( IBSP 179068 View Materials ) GoogleMaps   ; Cave E_056 (20°1'35''S; 43°41'8''W), 1F, 12–21.XI.2009 ( IBSP 179053 View Materials ) GoogleMaps   ; Cave E_071 (20°1'12''S; 43°42'48''W), 1F, 12–21.XI.2009 ( IBSP 179056 View Materials ) GoogleMaps   ; Cave E_072 (20°0'50''S; 43°43'31''W), 1F, 12– 21.XI.2009 ( IBSP 179064 View Materials ), 2F ( IBSP 179065 View Materials ), 1M, 1F ( IBSP 179066 View Materials ) all collected by R. Bessi et al. GoogleMaps   ; Rio Acima , Cave APOL_0024 (20°5'24''S; 43°41'9''W), 1F ( IBSP 179627 View Materials ) GoogleMaps   ; Cave SPC_040 (20°3'42''S; 43°40'15''W), 1F ( IBSP 179483 View Materials ) GoogleMaps   ; Cave SPD-033 (20°3'15''S; 43°39'56''W), 1F ( IBSP 179484 View Materials ) GoogleMaps   ; Cave SPC_035 (20°3'33''S; 43°40'13''W), 1F ( IBSP 179485 View Materials ), 1F ( IBSP 179489 View Materials ) GoogleMaps   ; Cave SPD-038 (20°3'42''S; 43°40'14''W), 3F ( IBSP 179490 View Materials ) all collected in between 25–27.VII.2011 by Equipe Carste coll. GoogleMaps   ; Santa Bárbara , Cave Caverna SG-002 (20°2'56''S; 43°40'55''W), 3F ( IBSP 179499 View Materials ) GoogleMaps   ; Cave Caverna SG-007 (20°2'58344''S; 43°41'5''W), 1F ( IBSP 179500 View Materials ), 2F ( IBSP 179501 View Materials )   ; Cave Caverna SG-009 (20°3'18''S; 43°41'9''W), 4F ( IBSP 179502 View Materials ), 2F ( IBSP 179503 View Materials ) all collected in between 26– 30.IX.2011 by Equipe Carste coll. GoogleMaps   ; Cave APOL_0007 (20°0'59''S; 43°39'57''W), 1F ( IBSP 179618 View Materials ) GoogleMaps   ; Cave APOL_ 0008 (20°1'9''S; 43°40'9''W), 1F ( IBSP 179619 View Materials ), 1F ( IBSP 179620 View Materials ) GoogleMaps   ; Cave APOL_0009 (20°1'15''S; 43°39'50''W), 2F ( IBSP 179621 View Materials ) GoogleMaps   ; Cave APOL_0013 (20°0'15''S; 43°39'51''W) 1F ( IBSP 179622 View Materials ) GoogleMaps   ; Cave APOL_0016 (20°3'11''S; 43°42'6''W), 2M14F ( IBSP 179623 View Materials ), 4F ( IBSP 179624 View Materials ) GoogleMaps   ; Cave APOL_0022 (20°3'56''S; 43°41'40''W), 2F ( IBSP 179625 View Materials ), 1F ( IBSP 179626 View Materials ) GoogleMaps   ; Cave APOL_0029 (20°2'35''S; 43°41'56''W), 2F ( IBSP 179628 View Materials ) GoogleMaps   ; Cave APOL_0030 (20°2'16116''S; 43°39'50''W), 1M, 2F ( IBSP 179629 View Materials )   ; Cave APOL_0001 (20°0'34''S; 43°39'56''W) 1F ( IBSP 179630 View Materials ) GoogleMaps   ; Cave APOL_0002 (20°0'36''S; 43°39'56''W), 1F ( IBSP 179631 View Materials ) GoogleMaps   ; Cave APOL_0005 (20°0'37''S; 43°39'47''W), 1F ( IBSP 179632 View Materials ) GoogleMaps   ; Cave APOL_0006 (20°0'59''S; 43°40'5''W), 1F ( IBSP 179633 View Materials ), 1F ( IBSP 179634 View Materials ) GoogleMaps   ; Cave APOL_0010 (19°59'57''S; 43°39'55''W), 4F ( IBSP 179635 View Materials ), 1F ( IBSP 179636 View Materials ), 3F ( IBSP 179637 View Materials ) GoogleMaps   ; Cave APOL_0012 (20°0'11''S; 43°40'8''W), 1F ( IBSP 179638 View Materials ) all collected in between 30.VI–15.VII.2011 by Equipe Carste coll. GoogleMaps   ; Conceição do Mato Dentro , Conceição Sul, Cave CAV-0012 (19°6'58''S; 43°25'0''W) 1F, 12–24.IX.2011 ( IBSP 180436), 2F, 13–17.II.2012 ( IBSP 180467) GoogleMaps   ; Cave CAV-0013 (19°7'5286''S; 43°26'0''W) 1F, 12–24.IX.2011 ( IBSP 180437), 1F, 13–17.II.2012 ( IBSP 180468)   ; Cave CAV-0014 (19°8'15''S; 43°24'19''W) 1F, 12–24.IX.2011 ( IBSP 180438), 1M ( IBSP 180439), 1F ( IBSP 180440), 1F ( IBSP 180441), 1F ( IBSP 180442), 1F ( IBSP 180443), 1F, 13–17.II.2012 ( IBSP 180469), 1F ( IBSP 180470), 1F ( IBSP 180471), 1F ( IBSP 180472) GoogleMaps   ; Cave Caverna CAV- 0015 (19°6462'0''S; 43°24'18''W), 1F, 13–17.II.2012 ( IBSP 180473)   ; Morro do Pilar : Cave CAV-0002 (19°9'20''S; 43°23'53''W) 2F, 12–24.IX.2011 ( IBSP 180427), 1F ( IBSP 180428), 1F, 13–17.II.2012 ( IBSP 180459), 2F ( IBSP 180460) GoogleMaps   ; Cave CAV-0003 (19°9'20''S; 43°23'52''W), 1F, 12–24.IX.2011 ( IBSP 180429), 1F, 13–17.II.2012 ( IBSP 180461) GoogleMaps   ; Cave CAV-0005 (19°14'58596''S; 43°22'6''W), 1F, 12–24.IX.2011 ( IBSP 180430), 1F ( IBSP 180431), 1F, 13–17.II.2012 ( IBSP 180462)   ; Cave CAV-0006 (19°15'56''S; 43°25'16''W), 1F, 12–24.IX.2011 ( IBSP 180432) GoogleMaps   ; Cave CAV-0009 (19°14'32''S; 43°22'2''W), 1F, 12–24.IX.2011 ( IBSP 180433), 1F ( IBSP 180434), 1F, 13–17.II.2012 ( IBSP 180463), 1F ( IBSP 180464) GoogleMaps   ; Cave CAV-0010 (19°9'27''S; 43°23'29''W), 1F, 12–24.IX.2011 ( IBSP 180435), 1F, 13–17.II.2012 ( IBSP 180465), 1F ( IBSP 180466) GoogleMaps   ; Cave CAV-0017 (19°12'4''S; 43°22'58''W), 1F, 12–24.IX.2011 ( IBSP 180444), 1F, 13–17.II.2012 ( IBSP 180474) GoogleMaps   ; Cave CAV-0018 (19°10'13''S; 43°23'46''W), 1F, 12–24.IX.2011 ( IBSP 180445), 1F, 13–17.II.2012 ( IBSP 180475), 1M, 2F ( IBSP 180476), 1M1F ( IBSP 180477) GoogleMaps   ; Cave CAV-0019 (19°9'44748''S; 43°23'29''W), 2F, 12–24.IX.2011 ( IBSP 180446), 2F, 13–17.II.2012 ( IBSP 180478), 1F ( IBSP 180479)   ; Cave CAV-0021 (19°8'55''S; 43°25'44''W), 1F, 12–24.IX.2011 ( IBSP 180447), 2F ( IBSP 180448), 1F, 13–17.II.2012 ( IBSP 180480), 1F ( IBSP 180481), 1F ( IBSP 180482), 1F ( IBSP 180483) GoogleMaps   ; Cave CAV-0022 (19°10'33''S; 43°24'42''W), 1F, 12–24.IX.2011 ( IBSP 180449) GoogleMaps   ; Cave CAV-0023 (19°13'15'' S; 43°23'26''W), 1F, 12–24.IX.2011 ( IBSP 180450), 1F, 13–17.II.2012 ( IBSP 180484), 1F ( IBSP 180485) GoogleMaps   ; Cave CAV-0001A (19°9'15''S; 43°24'13''W) 1F, 12–24.IX.2011 ( IBSP 180451), 1F ( IBSP 180453), 1F ( IBSP 180454), 1M, 1F, 28.II.2012 ( IBSP 180486), 1F ( IBSP 180487), 2F ( IBSP 180488), 1M ( IBSP 180489), 1F ( IBSP 180490), 1F ( IBSP 180491) GoogleMaps   ; Cave CAV-0001B (19°9'14''S; 43°24'10''W), 1F, 12–24.IX.2011 ( IBSP 180452), 1M, 1F ( IBSP 180455), 1F ( IBSP 180456), 1F ( IBSP 180457), 1F ( IBSP 180458), 1F, 28.II.2012 ( IBSP 180492), 1F ( IBSP 180493), 1F ( IBSP 180494) all colleted by R. Andrade & G. P. Perroni et al. GoogleMaps   ; Matozinhos: Cave MOC-66 (19°32'52''S; 44°0'37''W) 1 F, F. P. Franco & C. A. R. Souza et al. coll., 04–15.IV.2011 ( IBSP 180220). GoogleMaps  

Distribution. Brazil (caves in states of São Paulo and Minas Gerais, Fig. 32 View FIGURE 32 ).