Sagola denticollis Broun, 1880

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 45

Sagola denticollis Broun, 1880

( Figs. 21b, 21d , 22 )

Sagola denticollis Broun, 1880: 138 . Schaufuss 1888: 84; Raffray 1893: 33, 1904: 498, 1911: 6, 1924: 231; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.

Sagola elevata Broun, 1886: 886 . Raffray 1893: 38, 1904: 498. 1911: 6, 1924: 232; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.

Type Material Examined. Holotype of Sagola denticollis : NEW ZEALAND: Coromandel: 1♀ ( BMNH), glued on rectangular card, “253” [green label, printed]; “Type” [red label, printed]; “Tairua” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola denticollis .” [white label, handwritten] . Syntypes of Sagola elevata : NEW ZEALAND: Auckland: 1♀ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “1578.” [white label, handwritten]; “Woodhill” [white label, printed]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “ Sagola elevata .” [white label, handwritten]. 1♀ ( BMNH), glued on rectangular card, “1578.” [white label, handwritten]; “Woodhill” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] .

Additional Material (n = 18: 9 males; 9 females). NEW ZEALAND: Auckland : 1♂ 1♀, Bethells Matuku FS, 36′53S 174′28E, under bark #157, 29 I 1998, C. Carlton, R. Leschen ; 1♀, Waitakere R., Cascade-Kauri Park , up. kauri tr., 170m, 8 XII 1984 – 25 I 1985, kauri-podo-hdwd, A. Newton, M. Thayer 680, FIT & window trap ; 1♀, Matakana nr Warkworth , 27 XI 1995, J. Nunn, in decayed wood from bush remnant ; Coromandel: 5♂♂ , Little Barrier Island , 7 I 1952, rotten wood ; 2♀♀, Little Barrier Island , 6 I 1952, rotten wood ; 1♀, Great Barrier Island, Little Windy Hill , 13 XII 2002 – 17 I 2003, P. Sutton, forest edge malaise trap ; 1♀, Little Barrier Island , 17 XII 1983, J. Playfair ; Northland: 3♂♂ 2♀♀ (1♂, slide-mounted) , Parua , 4 XI 1981, G. Kuschel, litter and decayed wood 81/125 ;

Diagnosis. This species can be distinguished from S. parva by the transverse head, small eyes, enlarged antennomeres 4–6 of male with distinctive tubercles, and genitalia.

Redescription. Length 2.5–3.5 mm. Body brown, antennae, legs, and maxillary palpi paler, elytra yellowish brown ( Fig. 21b ). Head: Transverse, widest across eyes, frontal rostrum prominent ( Fig. 21b ). Male antennomeres 2–3 subquadrate, 4–6 enlarged, 6 largest, 7–10 subquadrate, 3–10 with distinctive tubercles. Female antennomeres 2–3 subquadrate, 4–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching hind point of eye from frontal fovea. Anterior frontal fovea oval, posterior frontal fovea surrounded by setae. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra subquadrate ( Fig. 21b ). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe with triangular lobe at midpoint ( Fig. 21d ). Phallobase of median lobe symmetrical and rectangular ( Fig. 21d ). Parameres asymmetrical, left paramere shorter than right, right divided anteriorly ( Fig. 21d ).

Type Locality. Tairua , Coromandel, New Zealand .

Distribution. Auckland, Coromandel, Northland ( Fig. 22 : triangles).

Habitat. Specimens of this species were collected using window and flight intercept traps or by sifting leaf litter or rotten wood in kauri, podocarp, and hardwood forests.

Comments. Specimens of S. denticollis are easily distinguished from other species externally by the transverse head with a prominent frontal rostrum. The type specimens of S. elevata share these diagnostic characters. For these reasons, we have placed S. elevata in synonymy with S. denticollis .

Figures

Fig. 21. Sagola species in the parva species-group. Habitus, dorsal views: a) S. parva, b) S. denticollis. Aedeagi, dorsal views: c) S. parva, d) S. denticollis. S. parva, male: e) Antenna, f) Dorsal surface of head, g) Ventral surface of head. Scale bars a–b = 1 mm, c–g = 0.1 mm.

Fig. 22. Known collection localities of New Zealand Sagola species. S. parva: black circles; S. denticollis: triangles.