Sagola bifida, Broun, 1915

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 24-25

3. bifida species-group

(11 species)

Diagnosis. The members of the bifida speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.4–3.2 mm; antennomere 1 approximately 2 times longer than wide with dull surface; ventral surface of male head depressed transversely, at least one-half covered with tubular structures on each side ( Fig. 12m ), with dense setae along edge; hind wings reduced to small pads; male fore femur with semicircular depression; male mid-femur with shallow depression ( Fig. 12n : arrow); male mid-tibia bent ( Fig. 12o : arrow); abdominal tergites IV–VI with discal carinae; present on North Island, not known from South Island ( Figs. 13–14 ).

KEY TO SPECIES OF THE BIFIDA SPECIES- GROUP

The key is based on genitalia because most specimens are indistinguishable based on external morphology.

1. Median lobe of genitalia deeply divided, major lobe broad and covering minor ( Fig. 12f )............. S. mayae new species

1′. Median lobe of genitalia not divided...2

2(1′). Median lobe of genitalia shorter than both parameres; right paramere twisted............................................................3

2′. Median lobe of genitalia longer than at least one of parameres; right paramere not twisted .............................................4

3(2). Apex of right paramere broad ( Fig. 12c ) .............................. S. crawi new species

3′. Apex of right paramere simple ( Fig. 12a ) ....................................... S. bifida Broun

4(2′). Right paramere with long minor branch ...............................................................5

4′. Right paramere without branch............7

5(4). Apical lobe of genitalia bent ventrally ( Fig. 12d ).............. S. earlyi new species

5′. Apical lobe of genitalia straight...........6

6(5′). Median lobe of genitalia cylindrical, longer than right paramere ( Fig. 12b ) .... ................................. S. latistriata Broun

6′. Median lobe of genitalia broad vertically, shorter than right paramere ( Fig. 12e ) .................. S. embersoni new species

7(4′). Left paramere with U-shaped depression apically ( Fig. 12g )................................... .................... S. ranatungae new species

7′. Left paramere without depression.........8

8(7′). Median lobe of genitalia shorter than left paramere, with simple apical lobe ( Fig. 12i )...... S. tekoauensis new species

8′. Median lobe of genitalia longer than parameres with modified apical lobe....9

9(8′). Apical lobe of genitalia broad with acute lobe behind apex ( Fig. 12h ) ................... ........................ S. baylessae new species

9′. Apical lobe of genitalia round and blunt .......................................................... 10

10(9′). Apical lobe of genitalia with semicircular depression; phallobase of genitalia transverse ( Fig. 12k ) ....................................... ........................ S. fairburni new species

10′. Apical lobe of genitalia with branch; phallobase of genitalia as long as wide ( Fig. 12j )............... S. ferroi new species

Figures

Fig. 12. Sagola species in the bifida species-group. Aedeagi, dorsal views: a) S. bifida, b) S. latistriata, c) S. crawi, d) S. earlyi, e) S. embersoni, f) S. mayae, g) S. ranatungae, h) S. baylessae, i) S. tekoauensis, j) S. ferroi, k) S. fairburni. S. latistriata, male: l) Dorsal surface of head, m) Ventral surface of head (arrow indicates large depression), n) Mid-femur, lateral view (arrow indicates shallow depression), o) Mid-tibia, lateral view (arrow indicates bend). Scale bars = 0.1 mm.

Fig. 13. Known collection localities of New Zealand Sagola species. S. bifida: black circles; S. latistriata: triangles; S. crawi: black square; S. earlyi: star; S. embersoni: white circle; S. mayae: white squares.

Fig. 14. Known collection localities of New Zealand Sagola species. S. ranatungae: black circle; S. baylessae: triangle; S. tekoauensis: black square; S. ferroi: star; S. fairburni: white circle.