Sagola laticeps Broun, 1911

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 136-137

Sagola laticeps Broun, 1911

( Figs. 68i , 69i , 70i , 72 )

Sagola laticeps Broun, 1911: 490 . Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.

Type Material. Holotype. NEW ZEALAND: Buller : 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “3362. ♂.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Greymouth.-Lewis-” [white label, handwritten]; “ Sagola ♂ laticeps.” [white label, handwritten]. The original label indicates the specimen is male, but it is female.

Additional Material (n = 29: 14 males; 15 females). NEW ZEALAND: Buller: 9♂♂ 5♀♀, Lewis Pass NR, 11.9km ese Springs Junction, 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 715, litter; 1♂ 1♀, Greymouth, Grandjeans tk, 10 XI 2005, R. Leschen, S. Nomura ; 1♂, 4km n of Rapahoe , 13 IX 1981, R. M. Emberson, coastal scrub and flax ; 1♀, Barrytown Croesus tk, 10 XI 2005, R. Leschen, S. Nomura, rotten log ; Westland: 1♂ 1♀ , Waiata Haast , 16 III 1968, R. A. Cumber, leaf litter ; 2♀♀, Otira, Barretts Creek , 9 II 1982, C.F. Butcher, litter ; 1♂ 2♀♀, Kelly Creek, Otira , 22 III 2009, J. Nunn, moss litter ; 1♂, Otira R, 6.8km ne Otira , 280m, 18–21 III 1980, Nothofagus fusca - podocarp, A. Newton, M. Thayer, litter ; 1♀, Doughboy Creek, 6km sw Mahitahi , 5 II 1984, J. C. Watt, wood mould ; 1♀, Lake Mahinapua , 20m, 23 X 1978, J.W. Early, broadleaf-punga litter ; 1♀, Arthur Pass NO, Otira , 580m, 1 III 1987, R. M. Emberson, rata-kamahi litter .

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.8–3.2 mm; temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.

Redescription. Length 2.8–3.2 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68i ). Head: Male head triangular ( Fig. 69i ), widest across temples. Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea round. Eye large and prominent, approximately one-half length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with flat ventral surface ( Fig. 69i ). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae ( Fig. 69i ). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular ( Fig. 68i ), female elytra subquadrate. Male meso-and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small, round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad, with pair of triangular processes apically ( Fig. 70i ). Phallobase of median lobe symmetrical and rounded ( Fig. 70i ). Parameres symmetrical and slender, with setae at apex and midpoint ( Fig. 70i ).

Type Locality. Greymouth , Buller, New Zealand .

Distribution. Buller, Westland ( Fig. 72 : black squares).

Habitat. Specimens of this species were collected by sifting leaf, wood, or moss litter in broadleaf or Nothofagus forests.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 72. Known collection localities of New Zealand Sagola species. S. pertinax: black circles; S. monticola: triangles; S. laticeps: black squares; S. castanea: stars; S. sheldoni: white circles; S. mercuryensis: white squares.