Sagola stewartensis Park and Carlton

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 145-146

Sagola stewartensis Park and Carlton , new species

( Figs. 68w , 69w , 70w , 74 )

Type Material. Holotype. NEW ZEALAND: Stewart Island: 1♂ ( NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ New Zealand: SI: Codfish IS nr summit, 244m, 14XII1966 J.I.Townsend, litter 66/443”, “HOLOTYPE Sagola stewartensis Park and Carlton 2013 ”. Paratypes (n = 24: 10 males; 14 females). NEW ZEALAND: Stewart Island: 1♂ 2♀♀ ( NZAC), Codfish I, North Hut tr, 6 XII 1981, S.A. Holloway, litter 81/207; 2♂♂ 3♀♀ ( NZAC), Pegasus Creek, 24 II 1968, G. Kuschel, litter 68/64; 5♂♂ 1♀ ( NZAC), Pegasus Creek, 24 II 1968, G. Kuschel, litter 68/63; 1♂ 1♀ ( NZAC), ne Big S. Cape I, 91m, 22 II 1969, J.S. Townsend, moss 69/83; 1♀ ( NZAC), Codfish I, Upper Valley tr, 27 XI 1981, B.A. Holloway, litter 81/197; 1♀ ( NZAC), Codfish I, Summit tk, 250m, 30 XI 1981, B.A. Holloway, litter 81/199; 1♀ ( NZAC), Codfish I, Upper Miro tr, 1 XII 1981, B.A. Holloway, litter 81/200; 1♀ ( NZAC), Garden Mound tr, 113m, 21 I 2007, R. Leschen, T. Buckley, K. Marske, log fogging; 1♂ 2♀♀ ( NZAC), Stewart Island, 14 II 1953, R. Hornabrook.

Etymology. This species is named after the type locality, Stewart Island.

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.6–2.9 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; ventral surface of male head with transverse, setose process behind mouthparts; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Stewart Island.

Description. Length 2.6–2.9 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68w ). Head: Male head triangular, widest across temples ( Fig. 69w ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface ( Fig. 69w ). Ventral surface of male head with transverse, setose process behind mouthparts ( Fig. 69w ). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate ( Fig. 68w ). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia broad with transverse apical lobe ( Fig. 70w ). Phallobase of median lobe symmetrical and rounded ( Fig. 70w ). Parameres symmetrical and slender, sparsely setose from midpoints to apices ( Fig. 70w ).

Distribution. Stewart Island ( Fig. 74 : white circles).

Habitat. Specimens of this species were collected mostly by sifting leaf and moss litter and one from log fogging.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 74. Known collection localities of New Zealand Sagola species. S. hanae: black circles; S. gyudongi: triangles; S. egmontensis: black squares; S. boonei: stars; S. stewartensis: white circles; S. jeongnamae: white squares.