Isthmohyla pictipes (Cope, 1876) FAIVOVICH & HADDAD & GARCIA & FROST & CAMPBELL & WHEELER, 2005

FAIVOVICH, JULIÁN, HADDAD, CÉLIO F. B., GARCIA, PAULO C. A., FROST, DARREL R., CAMPBELL, JONATHAN A. & WHEELER, WARD C., 2005, Systematic Review Of The Frog Family Hylidae, With Special Reference To Hylinae: Phylogenetic Analysis And Taxonomic Revision, Bulletin of the American Museum of Natural History 2005 (294), pp. 1-240 : 103

Isthmohyla pictipes Group

DIAGNOSIS: We are not aware of any synapomorphy supporting the monophyly of this group.

COMMENTS: We included a single exemplar of this group, and as such we did not test its monophyly. It is being tentatively recognized following Duellman (2001) until a rigorous test is performed. This species group is formed by the former Hyla lancasteri , H. pictipes , H. rivularis , and H. zeteki groups (Duellman, 1970, 2001). The monophyly of the former H. zeteki group does not seem to be controversial, as its two species share massive temporal musculature, bromeliad dwelling larvae, a terminal oral disc, and a labial tooth row formula of 1/1. The monophyly of the group composed of the former H. rivularis and H. pictipes groups (as defined by Duellman, 1970) is supported by the presence of an enlarged oral disc (Duellman, 2001) with a broad band of conic submarginal papillae on the posterior part of the disc, about three rows on the anterior part, and an M­shaped upper jaw sheath 27 (Faivovich, personal obs.; see also illustrations in Duellman, 2001). The monophyly of the former H. lancasteri group seems to be supported by the presence of granular dorsal skin (Duellman, 2001; known homoplastic instance in H. debilis ), a short snout, and the presence of dark ventral pigmentation (Wilson et al., 1994b, known homoplastic instance in H. thorectes .)

CONTENTS: Ten species. Isthmohyla calypsa (Lips, 1996) , new comb.; Isthmohyla debilis (Taylor, 1952) , new comb.; Isthmohyla insolita (McCranie, Wilson, and Williams, 1993) , new comb.; Isthmohyla lancasteri (Barbour, 1928) , new comb.; Isthmohyla picadoi (Dunn, 1937) , new comb.; Isthmohyla pictipes (Cope, 1876) , new comb.; Isthmohyla rivularis (Taylor, 1952) , new comb.; Isthmohyla tica (Starret, 1966) , new comb.; Isthmohyla xanthosticta (Duellman, 1968) ,

27 The description and illustrations of the tadpole of Hyla debilis by Duellman (1970) do not show these character states.

new comb.; Isthmohyla zeteki (Gaige, 1929) , new comb.