Dendropsophus marmoratus (Laurenti, 1768) FAIVOVICH & HADDAD & GARCIA & FROST & CAMPBELL & WHEELER, 2005

FAIVOVICH, JULIÁN, HADDAD, CÉLIO F. B., GARCIA, PAULO C. A., FROST, DARREL R., CAMPBELL, JONATHAN A. & WHEELER, WARD C., 2005, Systematic Review Of The Frog Family Hylidae, With Special Reference To Hylinae: Phylogenetic Analysis And Taxonomic Revision, Bulletin of the American Museum of Natural History 2005 (294), pp. 1-240 : 91

Dendropsophus marmoratus Group

DIAGNOSIS: This species group is diagnosed by 73 transformations in nuclear and mitochondrial protein and ribosomal genes. See appendix 5 for a complete list of these molecular synapomorphies. Possible morphological synapomorphies of this group are the warty skin around the margin of the low­ er lip, the crenulated margin of limbs, and the dorsal marbled pattern (Bokermann, 1964b) (instances of homoplasy in other Hylinae ). Furthermore, as it is discernible from the illustrations presented by Gomes and Peixoto (1991b) and Peixoto and Gomes (1999), and confirmed by Peixoto (personal commun. cited in Altig and McDiarmid, 1999a), known larvae of this species group share the presence of a thick sheath of tissue in the basal portion of the tail muscle and adjacent fins, another likely morphological synapomorphy.

COMMENTS: Bokermann (1964b) diagnosed this group as having large vocal sacs. While this could be a synapomorphy, we are hesitant to consider it as such until more anatomical and comparative studies are done in Dendropsophus . In connection with the large vocal sacs of the species of this group, Tyler (1971) mentioned that in Hyla marmorata the pectoral lymphatic septum is modified in a way that permits the inflated sac to intrude into sub­humeral spaces.

CONTENTS: Eight species. Dendropsophus acreanus (Bokermann, 1964) , new comb.; Dendropsophus dutrai (Gomes and Peixoto, 1996) , new comb.; Dendropsophus marmoratus (Laurenti, 1768) , new comb.; Dendropsophus melanargyreus (Cope, 1887) , new comb.; Dendropsophus nahdereri (B. Lutz and Bokermann, 1963) , new comb.; Dendropsophus novaisi (Bokermann, 1968) new comb.; Dendropsophus seniculus (Cope, 1868) , new comb.; Dendropsophus soaresi (Caramaschi and Jim, 1983) , new comb.