Hyloscirtus armatus (Boulenger, 1902) FAIVOVICH & HADDAD & GARCIA & FROST & CAMPBELL & WHEELER, 2005

FAIVOVICH, JULIÁN, HADDAD, CÉLIO F. B., GARCIA, PAULO C. A., FROST, DARREL R., CAMPBELL, JONATHAN A. & WHEELER, WARD C., 2005, Systematic Review Of The Frog Family Hylidae, With Special Reference To Hylinae: Phylogenetic Analysis And Taxonomic Revision, Bulletin of the American Museum of Natural History 2005 (294), pp. 1-240 : 84

Hyloscirtus armatus Group

DIAGNOSIS: This species group is diagnosed by 103 transformations in nuclear and mitochondrial protein and ribosomal genes. See appendix 5 for a complete list of these molecular synapomorphies. Duellman et al. (1997) suggested four synapomorphies of the H. armatus group: the presence of keratincovered bony spines on the proximal ventral surface of the humerus, on the expanded distal element of the prepollex, and on the first metacarpal; forearms hypertrophied; tadpole tail long with low fins and bluntly rounded tip; and the presence of a ‘‘shelf’’ on the larval upper jaw sheath.

COMMENTS: Our observations of breeding males of the two species of this group indicate the presence of darkly pigmented, keratinized spicules in the dorsum, head (particularly lips), forelimbs, undersides of forelimbs, and pectoral and abdominal region. As the breeding biology of this and the other two species groups of the genus becomes better known, it will be possible to understand if the presence of these spicules are a putative synapomorphy of the H. armatus group.

CONTENTS: Two species. Hyloscirtus armatus (Boulenger, 1902) , new comb.; Hyloscirtus charazani (Vellard, 1970) , new comb.