Pimpla rediviva Kopylov & Spasojevic & Klopfstein, 2018

Kopylov, D. S., Spasojevic, T. & Klopfstein, S., 2018, New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East, Zootaxa 4442 (2), pp. 319-330 : 325-326

Crusopimpla? rediviva ( Brues, 1910) , comb. nov.

Figure 4

* Pimpla rediviva: Brues 1910 , pp. 54–55, Fig. 40. Material. High-resolution photograph of the holotype examined (specimen #2161, Museum of Comparative Zoology , Harvard University, USA) . Florissant shales, Colorado, USA; Late Eocene. Imago, female in lateral aspect. Head, meso- and metasoma well preserved; antennae absent; one complete and one incomplete fore and one partial hind wing; partial hind legs; ovipositor and its sheaths probably in complete.

Redescription. Coloration. Head, mesosoma, hind coxa, and ventral part of hind femur dark brown; wing veins brown, light brown at base of pterostigma; metasomal tergites mostly reddish-brown, with dark areas basally.

Head. Head short and high, with eyes probably large.

Mesosoma. Mesosoma rather short and high, not well preserved and overlain by at least a fore wing, making interpretation of carinae somewhat difficult. Mesopleuron with epicnemical carina close to its front margin and probably meeting pronotum at mid-height; with a sinusoid line indicated which reaches from upper front corner to lower hind corner (might be an artefact). Metapleuron somewhat distorted, with pleural and submetapleural carinae complete. Propodeum with subcircular spiracle clearly shown, with complete lateral longitudinal carina, sublateral and probably also median portion of apical transverse carina and at least lateral portion of basal transverse carina; median part of propodeum not preserved, state of median longitudinal carinae unclear.

Wings. Fore wing 2.8× as long as wide; pterostigma 4.1× as long as wide; 1-Rs&1-M almost straight; 1mcu&2-Rs+M angled, but without any traces of 1-Rs+M; r-rs issues at mid-length of pterostigma, straight, 2.4× pterostigma width; areolet probably almost triangular, ~1.5× as long as wide, with 4-M very short; 2+3M 1.8× 2- Rs, 1.2× r-m. Hind wing not well preserved, veins unclear.

Legs. Legs stout, hind femur about 3.5x as long as wide.

Metasoma. Metasoma probably depressed on most of its length, but with apex somewhat compressed, almost twice as long as mesosoma. T1 mostly covered by hind leg, but lateroventral and laterodorsal carinae present at base; following tergites transverse, T7 not conspicuously shorter than T6. Ovipositor sheaths at least 1.35 mm or 0.25× metasomal length, with transverse wrinkles well visible; at least visible part of ovipositor straight, rather robust.

Measurements. Mesosoma length 2.6 mm, metasoma length 5.1 mm. Fore wing length 6.4 mm, width 2.3 mm.

Taxonomic placement. This species was placed in the genus Pimpla by Brues (1910), which at the time was interpreted in a much broader sense than today; the fossil has not been revised since. The lateral part of the propodeal carination is visible in this specimen, with the pleural, lateral longitudinal, and at least the sublateral portion of the apical transverse carinae present, forming part of the petiolar area. Unfortunately, the dorsal part of the propodeum is not preserved, but the correspondence of the fore wing venation, first metasomal tergite, robust ovipositor and thickened hind femur seem enough justification to place this species in the genus Crusopimpla (comb. nov.); as we cannot entirely exclude other genera, such as members of the Theronia genus group, we place it there with a question mark.

Figures

FIGURE 4. Crusopimpla? rediviva Brues holotype: a—part; b—our interpretation, where the wings were moved away so not to overlay the body. Dotted lines represent uncertain interpretations, grey lines reflect sculpture.