Homalocnemis nigripennis Philippi, 1865

Rafael, José A., Marques, Dayse W. A., González, Christian R. & Sinclair, Bradley J., 2022, Review of Homalocnemidae of Chile with a key to species and descriptions of male and female terminalia (Diptera: Empidoidea), Zootaxa 5222 (2), pp. 168-178 : 171-172

publication ID

https://doi.org/ 10.11646/zootaxa.5222.2.5

publication LSID

lsid:zoobank.org:pub:AB8A29BD-87BF-4913-8501-51AD7B647E67

DOI

https://doi.org/10.5281/zenodo.7469962

persistent identifier

https://treatment.plazi.org/id/03D9A938-AF1D-FF87-CE87-5BD7FD69A653

treatment provided by

Plazi

scientific name

Homalocnemis nigripennis Philippi, 1865
status

 

Homalocnemis nigripennis Philippi, 1865

( Figs 1–12 View FIGURES 1–5 View FIGURES 6–8 View FIGURES 9–12 , 28 View FIGURE 28 )

Homalocnemis nigripennis Philippi, 1865: 752 . Type-locality: Chile.

Holotype. Male (not found in MNNC, presumably lost). Chile.

Distribution. Chile: Valparaiso, Santiago ( Fig. 28 View FIGURE 28 ).

References. Reed, 1888: 301 (cat.); Bezzi, 1905: 457 (list); Kertész, 1909: 8 (cat.); Collin, 1928: 54 (com.); Melander, 1928: 14 (dist.); Collin, 1933: 29 (redes., fig. 6a); Stuardo, 1946: 104 (cat.); Smith, 1967: 9 (cat.); Chvála, 1991: 13 (com.); Camousseight, 2005: 90 (list); Yang et al., 2007: 280 (cat.).

Diagnosis. In addition to features in the key, this species is distinguished by the long, narrow digitiform epandrial process, male cerci separate from epandrium and female sternite 8 not greatly enlarged and apex weakly bilobed.

Redescription. Wing length: 5.0– 5.1 mm. Male ( Figs 1, 3 View FIGURES 1–5 , 6–8 View FIGURES 6–8 ). Predominantly gray pruinose, especially mesopleuron, postpronotal lobe and notopleuron; prescutellar depression with long, anteriorly pointed pruinescent stripe, extending to near apex of scutum; front of scutum gray pruinose, with brown vittae along dorsocentral rows, apex of scutum and postsutural region ( Figs 1, 4 View FIGURES 1–5 ). Palpus with long whitish setae. Wing darkly infuscate, anal lobe well developed in both sexes ( Figs 1–5 View FIGURES 1–5 ). Vein R 2+3 strongly upcurved distally around pterostigma, ending well proximal to level of branching of vein R 4+5. Section between veins R 2+3 and R 4 equal to or longer than section between veins R 4 and R 5. Hind tarsomere 1 slightly swollen ( Fig. 3 View FIGURES 1–5 ). Tergite 8 less than half-length of respective sternite ( Fig. 7 View FIGURES 6–8 ), narrowed medially ( Fig. 6 View FIGURES 6–8 ); sternite 8 with desclerotized or depigmented medial area, with intersegmental membrane pigmented or weakly sclerotized posterolaterally ( Fig. 7 View FIGURES 6–8 ); remaining pregenital sclerites unmodified. Terminalia not highly sclerotized. Epandrial lamellae subrectangular, fused narrowly along dorsal bridge; with narrow, digitiform subapical process; dorsal margin with long, dense setae, nearly as long as width of epandrium ( Fig. 6 View FIGURES 6–8 ). Cercus oval, rounded posteriorly, with short dense ventroapical setae; not fused to epandrium or surstylus ( Figs 6, 7 View FIGURES 6–8 ). Hypoproct membranous medially, clothed in setulae laterally. Subepandrial sclerite broadly V-shaped, extending as bacilliform sclerites to surstylus; surstylus arched dorsally with apical notch; basally fused with epandrium ( Figs 6, 8 View FIGURES 6–8 ). Hypandrium ( Figs 7, 8 View FIGURES 6–8 ) with gonocoxal apodeme short, narrow, not greatly expanded horizontally; gonocoxal apodemes united dorsally by upright phallic plate; phallic plate paired, narrowly extending dorsally to base of subepandrial sclerite, opposite dorsal bridge of epandrial lamellae. Postgonites not differentiated. Hypandrium mostly fused with phallus, with apex of phallus emerging apically ( Fig. 8 View FIGURES 6–8 ); without lateral projections. Ejaculatory apodeme slender, elongate, positioned within hypandrium ( Fig. 8 View FIGURES 6–8 ).

Female ( Figs 2, 4–5 View FIGURES 1–5 ). Same pattern of color as male ( Fig. 1 View FIGURES 1–5 ). Sternite 8 broadly joined ventrally, with apex weakly bilobed ( Figs 10, 11 View FIGURES 9–12 ), not greatly enlarged, subequal to anterior portion of sternite 8; sparsely clothed in short setae. Syntergite 9+10 well developed, completely divided medially ( Fig. 9 View FIGURES 9–12 ). Sternite 10 rectangular, weakly sclerotized; narrowly expanded along anterolateral margin of syntergite 9+10. Cercus ovoid fused to syntergite 9+10 ( Figs 9–11 View FIGURES 9–12 ). Spermatheca well sclerotized, tubular, tapered and sinuous apically ( Fig. 12 View FIGURES 9–12 ).

Material examined. CHILE. Santiago [= Valparaiso], Marga-Marga , 7.ix.1927, Jaffuel & Pirion (1 ♀, USNM) [33°02′57″S – 71°30′10″W]; GoogleMaps Province Santiago [Santiago, metropolis], Quebrada de la Plata , 27.x.1967, el. 510 m, 33°31′S – 70°47′W, E. Schlinger & M. Irwin (1♁, 1♀, CAS). GoogleMaps

Remarks. Marga Marga is located in the Valparaíso Region of the Province of Marga Marga, not Santiago Province as stated on the label of the studied specimen. As suggested by Collin (1933), H. nigripennis has a more northerly distribution than H. praesumpta ( Fig. 28 View FIGURE 28 ).

MNNC

Museo Nacional de Historia Natural, Santiago

USNM

Smithsonian Institution, National Museum of Natural History

CAS

California Academy of Sciences

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Homalocnemiidae

Genus

Homalocnemis

Loc

Homalocnemis nigripennis Philippi, 1865

Rafael, José A., Marques, Dayse W. A., González, Christian R. & Sinclair, Bradley J. 2022
2022
Loc

Homalocnemis nigripennis

Philippi, R. A. 1865: 752
1865
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