Manota spathula, Hippa, Heikki, 2007

Hippa, Heikki, 2007, The genus Manota Williston (Diptera: Mycetophilidae) in Melanesia and Oceania, Zootaxa 1502, pp. 1-44 : 19-21

Manota spathula sp. n.

( Figs 1 D, 2D, 10A–C)

A small-sized Manota .

Male. Coloration. Pale brown, frons, vertex and occiput darker brown, mouthparts and legs slightly paler than thorax but apex of coxa, trochanter and base and apex of femur on legs 2 and 3 darker brown, wing unicolorous, hyaline, pale yellowish-brown, knob of haltere black. Head. Antennal flagellomere 4, Fig. 10 C. Palpomere 3 with apicomesial extension, with ca. 4 apically expanded curved sensilla; palpomere 4 with parasegment. Thorax. Pleural setosity pattern, Fig. 1 D: anepisternum setose, with 21 setae; anterior basalare setose, with ca. 10 setae; preepisternum 2 setose, with 7 setae; laterotergite setose, with 18 setae; episternum 3 setose, with 8 setae. Wing. Length 1.6 mm. Hypopygium ( Figs 10 A, B). Sternite 9 laterally united with gonocoxa except for the extreme posterior part, posterior margin convex, at the level of the base of gonostylus, anterior margin shallowly notched, setosity similar to ventral setosity of gonocoxa. Parastylar lobe present, exposed but partly covered by the posterior part of sternite 9, short, associated with weakly sclerotized lobe in a more lateral position, with two setae directed obliquely posteriad. Paraapodemal lobe not exposed in ventral view of hypopygium. The dorsal mesial margin of gonocoxa with a large lobe at the middle, bearing many blunt-ended setae on the margin. Apicomesial angle of gonocoxa elongated lobe-like, covering the basal body of the juxtagonostylar megaseta. Number of juxtagonostylar setae one: it is a broad flattened megaseta arising from a basal body half the length of megaseta. Gonostylus elongate, widening towards the slightly bilobed apex, with two closely set very long apical setae, with a row of stronger dorsomesial setae diverging from the other rather weak setosity. Tegmen with weak lateral shoulders. Hypoproct long, extending posteriorly nearly as far as the apex of gonostylus, with ca. 15 scattered ventral setae on each half. Cerci separate.

Female. Similar to male. When seen in ventral view, antennal flagellomere 4 in both specimens with the length/width ratio 1.3–1.6. Palpomere 3 with 4–5 curved sensilla. Anepisternum with 30–49 setae, anterior basalare with 5–6 setae, preepisternum 2 with ca. 2–4 setae, laterotergite with 25–28 setae, episternum 3 with 11 setae. Wing length. 2.0– 2.3 mm. Apical part of abdomen, Fig. 2 D: the basal tubercles of the long setae on tergite 9 unusually small, as long as basally wide. Cercus two-segmented.

Preimaginal stages unknown.

Discussion. See under M. subspathula .

Types. Holotype. Male, Papua New Guinea, Morobe province, Ilauru, Mt. Kolorong, 2200 m, Malaise trap in forest close to field station, 26–30.X.1999, H. Hippa, R. Norberg and D. Borisch (in NRM).

Additional material. 1 female with same data as holotype (in NRM). 1 female with same data as holotype except for Malaise trap in forest near big lookout, 26–30.X.1999 (in NRM).

Figures

FIGURE 1. Pleura, lateral view. A. Manota serawei sp. n. (holotype). B. M. explicans sp. n. (paratype). C. M. perissochaeta sp. n. (paratype). D. M. spathula sp. n. (holotype). E. M. bicuspis sp. n. (paratype). Scale 0.10 mm.

FIGURE 2. Apical part of female abdomen, lateral view. A. Manota orthacantha sp. n. B. M. taedia Matile. C. M. ctenophora Matile. D. M. spathula sp. n. E. M. explicans sp. n. F. M. serawei sp. n. Scale 0.10 mm.

FIGURE 10. Manota spathula sp. n. (holotype): A. Hypopygium, dorsal view. B. Hypopygium, ventral view. C. Antennal flagellomere 4, lateral view. Scale for A and B 0.10 mm, for C 0.05 mm.