Mesembrinella guaramacalensis Whitworth

Whitworth, Terry L. & Yusseff-Vanegas, Sohath, 2019, A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea), Zootaxa 4659 (1), pp. 1-146: 50-54

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Mesembrinella guaramacalensis Whitworth

sp. nov.

Mesembrinella guaramacalensis Whitworth   , sp. nov.

( Figs 288 View FIGURES 288–293 , 332 View FIGURES 326–349 , 375 View FIGURES 370–400 , 417 View FIGURES 411–423 , 487–488 View FIGURES 484–487 View FIGURE 488 )

Diagnosis. A medium-sized fly averaging 10 mm (9–11/2) in length. Discs of upper and lower calypters light tan to brown, with brown fringe; anterior spiracle yellow vs. brown in M. spicata   ; T4 with marginal row of setae vs. without marginal row of setae in M. spicata   ; male unknown, female terminalia as in Fig. 288 View FIGURES 288–293 .

Description. Female. Frons 0.245 0.24–0.25/2 of head width at narrowest. Fronto-orbital yellow with pale yellow tomentum; frontal vitta dark orange below, brown above; parafacial mostly orange with yellow tomentum, posterior 1/4 dark with pale tomentum; gena with short horizontal row of 4 stout setae midway and scattering of fine dark setae overall; postgena dark silvery with fine yellow setae; occiput dark, like postgena, with silvery tomentum and fine yellow setae, median occipital sclerite shiny dark brown; antenna bright orange, arista orange at base, remainder tan; palpus typical; eye with median facets 2x size of lateral facets; ocellar triangle medium-sized, ocelli of equal size; facial ridge with only two dark supravibrissal setae and some small brown setae clustered at base.

Thorax without stripes on dorsum, bluish with heavy, irregular pale tomentum; pleura blue with pale tomentum; chaetotaxy: ac 2:1, dc 2:3, ia 0, ph 1, ppn 3x 3, kat 1:1, meral setae typical, 1 pair converging ap, sa absent, 1 lat, 1 stout bas, 1 pb, 1 disc; subscutellum moderately developed; anterior spiracle medium-sized, yellow, posterior spiracle medium-sized, brown; legs entirely brown. Wing hyaline; subcostal sclerite bare; basicosta tan, tegula brown; section IV 0.16 of section III; discs of upper and lower calypters brown; rim of upper calypter with short dark brown setae, rim of lower calypter with long dark brown setae.

Abdomen entirely purple with whitish tomentum; T4–5 with marginal rows of stout setae; T5 without discal setae, with only short fine setae. Terminalia. T6 of WU shape, T7 short and broad, recessed along rear edge midway, T8 as separate sclerites ( Fig. 288 View FIGURES 288–293 ); ST6–8 and hypoproct as in Fig. 332 View FIGURES 326–349 ; spermathecae filiform ( Fig. 375 View FIGURES 370–400 ); ST1–5 as in Fig. 417 View FIGURES 411–423 .

Male. Unknown.

Type material. HOLOTYPE ♀ ♦* ( TLW294) ( Venezuela , Trujillo; MJMO; Fig. 487 View FIGURES 484–487 ), labeled: VENEZUELA Trujillo / P.N. Guaramacal 1480 m / 11–16 /VI/2002 / 9°19ʹ021 N– 70°15ʹ480 W; T. Amarilla / R. Briceño; J. Clavijo; R. Paz, F. Díaz; L. Joly; A. / Chacón / Proyecto S 1-2000000479   ; HOLOTYPE / Mesembrinella   / guaramacalensis / T.L. Whitworth.

ALLOTYPE ♀ ♦ ( TLW293): same data as holotype except 14–20.ii.2002 ( MJMO).  

Distribution. Venezuela.

Remarks. Known only from two females from Venezuela. The two female specimens ( TLW 293–294) were barcoded and the sequence data place them in the spicata   group, in a distinct cluster near M. mexicana   sp. nov. ( Fig. 488 View FIGURE 488 ). Barcodes show that this species has a 4% genetic distance from M. mexicana   and an 8% genetic distance from M. spicata   .

Etymology. The name guaramacalensis   refers to the type locality of the new species, Guaramacal National Park in Venezuela.


Wayland University


Universidad Centro Occidental, Decanato de Agronomia