Alloxysta pilipennis ( Hartig, 1840 )

Ferrer-Suay, Mar, Selfa, Jesús & Pujade-Villar, Juli, 2018, Palaearctic species of Charipinae (Hymenoptera, Figitidae): two new species, synthesis and identification key, European Journal of Taxonomy 427, pp. 1-110 : 63-64

Alloxysta pilipennis ( Hartig, 1840)

Figs 3.3, 5.7

Xystus pilipennis Hartig, 1840: 199 . Type: deposited in ZSM (examined).

Allotria pilipennis – Thomson 1862: 406.

Allotria (Allotria) pilipennis – Dalla Torre & Kieffer 1902: 40.

Charips (Charips) pilipennis – Dalla Torre & Kieffer 1910: 283.

Alloxysta pilipennis – Hellén 1963: 19.

Diagnosis

Alloxysta pilipennis is mainly characterized by a closed radial cell that is 2.5 times as long as wide (Fig. 5.7); the presence of pronotal and propodeal carinae; and female antenna with rhinaria beginning on F3, F1 longer than pedicel and F2, and F2–F4 subequal in length (Fig. 3.3). It is similar to A. pusilla , but can be differentiated by the proportions of the flagellomere in females (F2 is subequal to F 3 in A. pilipennis females (Fig. 3.3), but shorter than F 3 in A. pusilla females (Fig. 3.7)), flagellomere proportions and shape in males (F1–F3 are not subequal and not curved in A. pilipennis males, but F1–F3 are subequal in length and slightly curved in A. pusilla males), and the size of the radial cell (2.5 times as long as wide in A. pilipennis females (Fig. 5.7), but 2.7 times in A. pusilla females (Fig. 5.11)).

Material examined

Lectotype

GERMANY: ♀ [1709 (blue label)], [♀], [lectotype H.H. Evenhuis (orange label)], [ Xystus pilipennis Hartig det. H.H. Evenhuis 1980], [ Alloxysta pilipennis ( Hartig, 1840) ♀ M. Ferrer-Suay det. 2011] ( ZSM).

Paralectotypes

GERMANY: 1 ♀ [711 (red label)], [Paralectotype Xystus pilipennis Hartig, 1840 ♀ (red label)], [ Alloxysta pilipennis ( Hartig, 1840) ♀ M. Ferrer-Suay det. 2011] ( ZSM); 4 ♀♀ [Paralectotype Xystus pilipennis Hartig, 1840 ♀ (red label)], [ Alloxysta pilipennis ( Hartig, 1840) ♀ M. Ferrer-Suay det. 2011] ( ZSM).

Additional specimens (11 ♂♂, 41 ♀♀)

CZECH REPUBLIC: 1 ♀ [Moravia, Palava near Mikulov, = 400 m, 9 Aug. 1991, L. Masner, sweep] ( CNCI C-264); 1 ♀ [Moravia, Ladnice environs, 7–9 Aug. 1991, L. Masner riparian, forest] ( CNCI C-272).

GERMANY: 1 ♀ [Hesse, Gelnhausen, Sep. 1967, MT,A.W. Steffan] ( CNCI C-228); 5 ♀♀ [Schwarzwald, Todtmoos environs, 6 Aug. 1984, L. Masner, screen sweeping] ( CNCI C-235).

IRAN: 1 ♀ [Damavand, 40 km, E. of Tehran in orchard, 7–15 Jul. 1977, M. Cox] ( BMNH). 1 ♀ [Tehran Prov., Shahdasht, 25 Jun. 1978, J.T. Huber] ( CNCI C-317); 1 ♂, 1 ♀ [Karaj, 40 km W Tehran, 25–28 Jun. 1978, J.T. Huber, YPT] ( CNCI C-316).

JAPAN: 2 ♀♀ [Aichi Shitara, Uradani, 900 m, beech forest, 23–29 May 1994, K. Yamagishi, YPT] ( CNCI C-187); 1 ♀ [Iwate, Mt. Hayachine, 21 Jun. 1989, M.J. Sharkey, sweep] ( CNCI C-198); 1 ♀ [Ibaraki, Tsukuba, NIAES, 14–21 Jul. 1989, M.J. Sharkey, PT] ( CNCI C-191); 1 ♀ [Kyushu, 700 m, Fukuoka, MT. Hiko, 10–21 Jul. 1989, MT, K. Takeno and M. Sharkey] ( CNCI C-171); 1 ♀ [Hokkaido, Aizan, 800 m, 4 Jul. 1989, sweep, M.J. Sharkey] ( CNCI C-186); 1 ♀ [Fukuoka, Mt. Hiko, 10–21 Jul. 1989, Takeno and M. Sharkey] ( CNCI C-154); 1 ♀ [Fukuoka, Mt. Hiko, 18–25 Sep. 1989, K. Takeno and M. Sharkey] ( CNCI C-159); 1 ♀ [Hokkaido, Furano, Exp. Forest, 43°15′ N, 142°20′ E, 9 Aug. 1996, 500 m, L. Masner, sweep] ( CNCI C-166); 1 ♀ [Fukuoka, Mt. Hiko, 12–29 May 1989, Takeno and Sharkey] ( CNCI C-145); 1 ♀ [Fukuoka, Mt. Hiko, 700 m, 28 Apr.–10 May 1989, M.J. Sharkey, MT] ( CNCI C-139); 1 ♂ [Hokkaido, Hidako Mts below Pyo Tan, 500 m, 14 Aug. 1996, L. Masner, S.S. J-12] ( CNCI C-161); 1 ♀ [Aichi, Midori -Ku, Nagoya city, 2 Jun. 1984, R. Baczyuski] ( CNCI C-180); 1 ♂ [Hokkaido, Furano, Exp. Forest, 43°15′ N, 142°20′ E, 9 Aug. 1996, 500 m, L. Masner, sweep] ( CNCI C-163); 1 ♀ [Aichi, Shitara, Uradani, 900 m, 13–19 May 1994, K. Yamagishi, EMT, beech forest] ( CNCI C-223); 2 ♀♀ [Hokkaido, Sapporo, Jozankei, 350 m, 10–21 Aug. 1989, K. Maeto and M. Sharkey, MT] ( CNCI C-206); 3 ♀♀ [Aichi, Shitara, Uradani, 900 m, 2–8 May 1994, K. Yamagishi, EMT, beech forest] ( CNCI C-224); 2 ♀♀ [Iwate, Mt. Hayachine, 21 Jun. 1989, M. Sharkey, sweep] ( CNCI C-208); 2 ♀♀ [Aichi, Shitara, Uradani, 900 m, 4–10 Jul. 1994, K. Yamagishi, EMT, beech forest] ( CNCI C-217).

RUSSIA: 2 ♀♀ [Russian Far East, Primorski Krai Lazovski Zapovednik, c.170 Km, E. Vladivostok, Ta-Chingousa, 43°00′42″ N 134°07′34″ E, Om 28 Aug.–16 Sep. 2001, sandy coast, Malaise trap 494°, M. Quest coll., Alloxysta spp-indet dwarfs and intermediates] ( BMNH B.M. 2009-59); 1 ♂ [Primoskiy Kray Ussyriysk District, Gomotayozhnoye, 11–15 Aug. 2003, Malaise trap, M.V. Michailovskaya] ( USNM); 1 ♀ [Primorskiy Kray, Vladivostok environs, ??. ??.1992, A. Okulov] ( CNCI C-297); 1 ♀ [Magadanskaya Oblast, Aborigen Field Station, 500 m, 25 Jul.–10 Aug. 1990, D.M. Wood] ( CNCI C-298).

SWEDEN: 1 ♂ [Sk., Lund Zoological Mus, grounds, Aug. 1976] ( BMNH).

SWITZERLAND: 6 ♂♂, 3 ♀♀ [Dielsdorf, 650 m, 17 Aug. 1984, L. Masner, sweep] ( CNCI C-290).

UNITED KINGDOM: England: 1 ♀ [Surrey, Box Hill (near Dorking) 28 Aug. 1982, E.E. Grissell herbaceous vegetation] ( USNM).

Distribution

Holarctic and Neotropical.

Certain records: Colombia ( Ferrer-Suay et al. 2012f: 325), Germany ( Hartig 1841: 352), Iran (Ferrer- Suay et al. 2013a: 39), Spain ( Ferrer-Suay et al. 2013g: 327).

Uncertain records: Austria ( Giraud 1860: 129; Hellén 1963: 16), England ( Dalla Torre & Kieffer 1910: 282), Finland ( Hellén 1963: 16), France ( De Gaulle 1908: 26; Dalla Torre & Kieffer 1910: 282), Israel ( Argaman 1988: 115), Poland ( Kierych 1979b: 16), Norway ( Hellén 1966: 393), Romania ( Ionescu 1969: 249), Scotland ( Cameron 1886: 85), Sweden ( Thomson 1862: 407), Switzerland ( Hellén 1963: 16), The Netherlands ( Hellén 1963: 16), USA ( Andrews 1978: 61).

New records: Czech Republic, Iran, Japan and Russia.