Innobindus oppositus, Löcker, 2023

Löcker, Birgit, 2023, Six new species in the genus Innobindus Jacobi 1928 (Hemiptera: Auchenorrhyncha: Fulgoromorpha: Cixiidae: Brixiini), European Journal of Taxonomy 908, pp. 108-134 : 126-130

Innobindus oppositus sp. nov.

urn:lsid:zoobank.org:act:27B9B898-05EB-433E-B2DB-5AA5A11D3DD9

Figs 7 , 13 , 14A

Diagnosis

Innobindus oppositus sp. nov. can be distinguished from all other species of Innobindus by the presence of 7 apical spines on the hind tibia. The forewing colouration of I. oppositus is also unique within Innobindus ; no other species has a dark longitudinal band along the anterior margin of the wing (formed by an entirely dark postcostal cell and dark pterostigma, and only interrupted by a pale stripe at base of pterostigma) ( Fig. 7A ). All other species either have an entirely colourless postcostal cell, e.g., I. gimani ( Fig. 5A ) or have alternating pale and dark sections in the postcostal cell, e.g., I. artus ( Fig. 2A ), I. loriensis ( Fig. 4A ) and I. kaanti ( Fig. 3A ).

Etymology

The Latin term ‘ oppositus ’ means ‘set against’ or ‘opposite’. Named after the arrangement of the aedeagal spines in left lateral view which are set against each other with their tips pointing in opposite directions.

Material examined

Holotype AUSTRALIA – NSW • ♂; Dorrigo Nat. Pk. ; [30.362º S, 152.731º E]; 11 Nov. 1961; C.W. Frazier leg.; ASCU ASCTHE017104 .

Description

COLOUR. Central parts of frons and clypeus light brown, lateral parts whitish. Anteclypeus mid to dark brown. Vertex mid to dark brown. Pronotum whitish apart from darker anterior parts. Mesonotum mid brown. Forewings hyaline colourless except for a mid to dark brown longitudinal band along the anterior margin of the wing (formed by dark postcostal cell and dark pterostigma and only interrupted by a pale stripe at base of pterostigma), a light to mid brown transverse band at level of pterostigma; veins light to dark brown; tubercles concolorous with veins. Legs light brown. Body dark brown.

MEASUREMENTS. Body length: ♂ 6.2 mm.

HEAD. Vertex about 1.1 × as long as wide; apical transverse carina incised (u-shaped); subapical transverse carina slightly u-shaped; median carina of vertex covering about ⅓ of entire length of basal compartment. Frons about 2.3 × longer than wide at frontoclypeal suture, about 2.0× longer in widest part; maximum width of frons more than twice apical width, steadily broadening; frons near apical transverse carina very narrow (about 3× as wide as median ocellus of frons); lateral carinae of frons in facial view convex, rectilinear apically; foliaceous, slightly extended laterally.

THORAX. Hind margin of pronotum more or less rectangular. Forewing about 3.1× longer than wide; costal margin with about 4–5 very indistinct tubercles; fork of ScP+RA and RP at same level as fork CuA1 and CuA2; crossvein r–m 1 slightly basad of, or at same level as fork MP1+2 and MP3+4; transverse veinlet m–cu 1 where it inserts at CuA distinctly basad of transverse veinlet r–m 1; transverse veinlet m–cu 1 where it inserts at MP3+4 at same level as transverse veinlet r–m 1; icu, where it inserts at CuA, distinctly distad of apex of clavus; RA unforked; MP3+4 bifid; crossvein m–cu 2 (delimiting subapical cell C4) distinctly distad of crossvein icua (delimiting subapical cell C5); nodus of y-vein slightly distad of centre of clavus. Hind leg: tibia with 7 apical spines and 6–7 very small to small lateral spines; 1 st tarsomere with 7 apical teeth and no platellae; 2 nd tarsomere with 8 apical teeth, no platellae, but up to 3 very fine setae.

MALE GENITALIA. Anal tube as in Fig. 13C–D . Pygofer and genital styles as in Fig. 13E–F : ventromedian process of pygofer narrow, subtriangular. Shaft of genital styles without indentation.Aedeagus ( Fig. 13A– B ): phallotheca left laterally with a long spine (a), bent about 90 degrees in distal half; ventrally with a long, curved spine (b); dorso-(right)laterally with a very long, very thin spine (c), covering almost entire length of phallotheca. Flagellum without spines or spine-like ridges.

Distribution

Central NSW ( Fig. 14A ).

Figures

Fig. 7. Innobindus oppositus sp. nov., holotype, ♂ (ASCU ASCTHE017104). A. Habitus. B–D, F. Head and thorax. E. Hindwing.

Fig. 13. Innobindus oppositus sp. nov., holotype, ♂ (ASCU ASCTHE017104). A. Aedeagus, left lateral view. B. Aedeagus, right lateral view. C–D. Anal tube. E–F. Genital styles.

Fig. 14. Known distribution of species of Innobindus Jacobi, 1928. A. artus group and I. oppositus sp. nov. B. licinus group. C. multimaculatus group.

Fig. 5. Innobindus gimani sp. nov., holotype, ♂ (ASCU ASCT00181701). A. Habitus. B–D. Head and thorax.

Fig. 2. Innobindus artus sp. nov., holotype, ♂ (QM T258320). A. Habitus. B–D. Head and thorax.

Fig. 4. Innobindus loriensis sp. nov., holotype, ♂ (ASCU ASCTHE017059). A. Habitus. B–D. Head and thorax.

Fig. 3. Innobindus kaanti sp. nov., paratype, ♂ (ASCU ASCTHE029135). A. Habitus. B–D. Head and thorax.