Amphidromus (Amphidromus) protania Lehmann & Maassen, 2004

Inkhavilay, Khamla, Sutcharit, Chirasak & Panha, Somsak, 2017, Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species, European Journal of Taxonomy 330, pp. 1-40 : 15

Amphidromus (Amphidromus) protania Lehmann & Maassen, 2004

Figs 1 , 4H

Amphidromus (Amphidromus) protania Lehmann & Maassen, 2004: 17–20 , figs 1–4. Type locality: South Laos, Salavan Province, near Ban Donxé Village, E bank of Se Don River.

Material examined

LAOS: Dextral, holotype ( RMNH 98143, Fig. 4 H ); 4D shells, paratypes ( RMNH 98144).

Remarks

No specimen was found in this survey. Amphidromus protania has a very similar shell shape to that the dipterocarp forest species A. givenchyi . The genitalia contain a long and coiled proximal part of the epiphallus, with a relatively short appendix, as in A. givenchyi ( Lehmann & Maassen 2004; Sutcharit & Panha 2006). However, the characters distinguishing it from A. givenchyi are the relatively smaller shell (height 26–31 mm in A. protania ; Table 1 ), the yellowish shell color as well as the brownish subsutural and spiral bands on the periphery. Moreover, Lehmann & Maassen (2004) mentioned that the brownish banding pattern had less variation. However, a brownish spiral band on the periphery can also be observed in juvenile A. givenchyi (see Sutcharit & Panha 2006: fig. 4o). Moreover, this species occurs in dipterocarp forest within the range of A. givenchyi ; therefore, it is possibly a shell color variant of A. givenchyi . Additional specimens for further anatomical and molecular study are desirable to test this hypothesis.

Figures

Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study.

Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann & Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020).