Copestylum quadratum, (WILLISTON, 1891)

Rotheray, Graham E., Marcos-García, Maria-Angeles, Hancock, Geoff, Pérez-Bañón, Celeste & Maier, Chris T., 2009, Neotropical Copestylum (Diptera, Syrphidae) breeding in Agavaceae and Cactaceae including seven new species, Zoological Journal of the Linnean Society 156 (4), pp. 697-749 : 717

COPESTYLUM QUADRATUM (WILLISTON, 1891)

Larva and puparium: Anterior fold with band of spicules narrowing towards the lateral ends. Longest vestiture of dorsum of prothorax longer than longest spicules on the anterior fold. Posterior breathing tube with AP about as long as wide above inconspicuous TR. AP tapering to about two-thirds of width of TR and 1.1¥ as long as TR is wide. Pupal spiracles about the same length as the AP and separated by about 1.3¥ their length with openings surrounding the apex.

Taxonomic notes: Copestylum quadratum is most similar to C. milae from which it differs by having the openings of the pupal spiracles surrounding the apex rather than being on the apex and by having the AP slightly longer than the TR is wide. Copestylum quadratum is separated from C. hidalgense by the absence of spicules on the anterodorsal margin of the mesothorax.

Material examined (INBio): 1 ♂, one puparium COSTA RICA San José, Santa Cruz, Central 5.ix.1999 Y. Cárdenas ex decaying cactus; 1 ♂, one puparium COSTA RICA Alajuela, Arado 4.xii.1999 Y. Cárdenas ex decaying cactus; 2 ♂, 1 ♀, three puparia, four larvae COSTA RICA Guanacaste, La Cruz, Santa Elena, Est. Junquilla 29.x.1999 Y. Cárdenas and D. Briceño ex decaying Stenocereus aragonii (F.A.C. Weber) Buxb. 1961 (Cactaceae) .