Nonveilleridia, Lelej, 1996

Williams, Kevin A., Lelej, Arkady S., Okayasu, Juriya, Borkent, Christopher J., Malee, Rufeah, Thoawan, Kodeeyah & Thaochan, Narit, 2019, The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia, Zootaxa 4602 (1), pp. 1-69 : 6-8

publication ID	https://doi.org/ 10.5281/zenodo.2669927
publication LSID	lsid:zoobank.org:pub:F066A962-743F-4899-AFAE-485C5A51EF2F
persistent identifier	https://treatment.plazi.org/id/03DCF504-FFF0-9878-FF39-28344CFBC493
treatment provided by	Plazi
scientific name	Nonveilleridia
status

Treatment

Key to southeastern Oriental mutillid genera: females (unknown in Nonveilleridia )

1. Eye setose, sometimes the setae rare and visible only under high magnification (Subfamily Ticoplinae ; one genus in SE Asia)................................................................ 1. Eosmicromyrmilla Lelej & Krombein, 2001

- Eye not setose........................................................................................ 2

2. Antenna short, club-like ( Fig. 1 View FIGURES 1–28 ); second metasomal segment lacking felt lines ( Fig. 30 View FIGURES 29–35 ); maxillary and labial palps number 2+2 ( Rhopalomutillinae ; one genus in SE Asia)........................................ 2. Pherotilla Brothers, 2015

- Antenna longer, more slender ( Figs 2–5 View FIGURES 1–28 ); T2 or S2 with felt line; maxillary and labial palps number 6+4............... 3

3. Mesopleuron more or less convex, mesonotum wider than pronotum or propodeum ( Figs 2, 18, 19 View FIGURES 1–28 ).................... 4

- Mesopleuron concave, mesoscutellum not wider than pronotum or propodeum ( Figs 3–6 View FIGURES 1–28 )............................ 8

4. T1 petiolate ( Fig. 2 View FIGURES 1–28 ); felt line present on S2 (Dasylabrinae; one genus in SE Asia)............... 3. Orientilla Lelej, 1979

- T1 wide, sessile or disciform; felt line present on T2 ( Mutillinae : Mutillini : Ephutina)............................... 5

5. Posterior border of T2 with wide shiny cuticular band, not punctured; head with long thickened setae ( Figs 15, 16 View FIGURES 1–28 )........ 6

- Posterior border of T2 with usual punctures; head with simple setae ( Figs 18, 19 View FIGURES 1–28 )...................................7

6. Head width 1.2 × mesosoma width ( Fig. 15 View FIGURES 1–28 ); compound eye flat, small; maxillary palps approximately as long as scape........................................................................ 16. Cockerellidia Lelej & Krombein, 1999

- Head width 0.9 × mesosoma width ( Fig. 16 View FIGURES 1–28 ); compound eye convex, usual; maxillary palps much longer than scape........................................................................ 17. Karlidia Lelej in Lelej & Krombein, 1999

7. T2 with large deep antero-lateral pubescent fovea................................ 18. Odontomutilla Ashmead, 1899

- T2 without antero-lateral pubescent fovea, with linear felt line........................ 19. Yamanetilla Lelej, 1996

8. Mesopleural suture ending at mesonotal tubercle, complete to dorsum of mesosoma ( Figs 37, 46 View FIGURES 36–47 ); inner margin of mandible with distinct tooth in basal third ( Figs 38, 41 View FIGURES 36–47 ) ( Myrmillinae ). One genus in SE Asia........... 4. Bischoffitilla Lelej, 2002

- Mesopleural suture ending near anterior spiracle or becoming obsolete before dorsum of mesosoma ( Figs 60 View FIGURES 56–67 , 75 View FIGURES 68–76 , 85 View FIGURES 77–85 ); inner margin of mandible usually with teeth only near apex ( Figs 133 View FIGURES 125–136 , 149 View FIGURES 147–154 ) ( Mutillinae ).................................. 9

9. Propodeum posterad with transverse row of denticles ( Figs 48, 52 View FIGURES 48–55 ) ( Mutillinae : Ctenotillini )......................... 10

- Propodeum posterad without transverse row of denticles ( Figs 59 View FIGURES 56–67 , 72 View FIGURES 68–76 )........................................... 11

10. T2 without pale spots in basal half ( Fig. 4 View FIGURES 1–28 ); pygidium microreticulate ( Fig. 51 View FIGURES 48–55 ); inner margin of mandible without subbasal denticle...................................................................... 5. Ctenotilla Bischoff, 1920

- T2 with two pale spots located transversely near the middle ( Fig. 5 View FIGURES 1–28 ); pygidium mostly smooth ( Fig. 55 View FIGURES 48–55 ); inner margin of mandible with small subbasal denticle................................................. 6. Zeugomutilla Chen, 1957

11. T1 very wide, slightly narrower than T2. ( Mutillinae : Mutillini : Mutillina)....................................... 12

- T1 not so wide, much narrower than T2................................................................... 13

12. T2 disc without any band or spots.................................................... 21. Mutilla Linnaeus, 1758

- T2 disc with two ovate pale spots disposed transversely................................ 20. Kurzenkotilla Lelej, 2005

13. T2 disc with one or three basal spots of pale setae disposed transversely ( Figs 8–12, 14 View FIGURES 1–28 ), without spot of dense setae ( Figs 6, 7, 13 View FIGURES 1–28 ), or with two small spots at the lateral extremes of T2 ( Figs 78 View FIGURES 77–85 , 90) ( Mutillinae : Smicromyrmini )............... 14

- T2 with two basal spots of pale setae disposed transversely ( Figs 20–28 View FIGURES 1–28 ) ( Mutillinae : Trogaspidiini).............. 23

14. T6 convex, without distinct pygidial area, with glabrous shiny area widened posterad, at most its apical part weakly carinated laterally............................................................................................ 15

- T6 flattened, with distinct pygidial area carinated laterally, pygidial area longitudinally striate, granulose or with glabrous shining apical part or entirely smooth, shiny................................................................... 18

15. Mesosoma distinctly evenly narrowed posteriorly ( Fig. 65 View FIGURES 56–67 )............................... 13. Promecilla André, 1902

- Mesosoma slightly broadened posteriorly or more or less parallel-sided ( Figs 56, 59 View FIGURES 56–67 , 68 View FIGURES 68–76 )............................ 16

16. Scutellar scal totally lacking........................................................ 11. Nordeniella Lelej, 2005

- Scutellar scale incospicuous, but distinct.................................................................. 17

17. T6 with glabrous shiny medial part, not carinated laterad even apically, considerably convergent basally where with lateral tuft of dense pale setae.................................................................. 14. Sinotilla Lelej, 1995

- T6 with elongate shiny glabrous pygidial area weakly carinated laterad........................ 8. Ephucilla Lelej, 1995

18. Scutellar scale lacking; mandible wide, tridentate apically............................... 7. Andreimyrme Lelej, 1995

- Scutellar scale more or less developed; mandible narrower.................................................... 19

19. Pygidial area strongly narrowed basally; T2 with one basal, one apical and two lateral pale spots ( Fig. 11 View FIGURES 1–28 )........................................................................................... 12. Physetopoda Schuster, 1949

- Pygidial area with more or less parallel lateral sides or widened basally. T2 with one or three pale basal spots and pale apical band ( Figs 8, 9 View FIGURES 1–28 ), if sometimes with apical medial spot presented, then lateral spots absent........................... 20

20. Pygidial area short oval ( Fig. 97 View FIGURES 95–106 ); T2 apical band medially expanded to apparent white setal spot ( Fig. 95 View FIGURES 95–106 )...................................................................................................10. Nemka Lelej, 1985

- Pygidial area elongated, with parallel sides or widened basally ( Figs 81 View FIGURES 77–85 , 88, 100, 106); T2 apical band not so expanded.... 21

21. Scutellar scale more than 2 times wider than its length; T2 with three basal setal spots disposed in horizontal line ( Figs 82, 84 View FIGURES 77–85 , 89, 92) or with two small spots at lateral extremes of T2 ( Figs 78 View FIGURES 77–85 , 90) and at most with pale median apical patch ( Fig. 84 View FIGURES 77–85 )............................................................................ 9. Mickelomyrme Lelej, 1995, part.

- Scutellar scale slightly wider than its length; T2 with lateral pale spots (if present) disposed behind median spot and with pale apical band or apical fringe ( Figs 107, 111 View FIGURES 107–114 )................................................................ 22

22. Longitudinal eye diameter 1.5–2.0 × distance between posterior eye border and posterior head border.............................................................................................. 15. Smicromyrme Thomson, 1870

- Longitudinal eye diameter at least 3.0 × distance between posterior eye border and posterior head border...................................................................................... Mickelomyrme chinensis ( Smith, 1855)

23. T6 without any pygidial area, convex, smooth, shiny, punctured basally or throughout.............................. 24

- T6 with more or less developed pygidial area, at least flattened shiny area carinated laterally......................... 25

24. Scutellar scale small but visible ( Figs 137–140 View FIGURES 137–146 ).......................................... 25. Orientidia Lelej, 1996

- Scutellar scale lacking ( Figs 145, 146 View FIGURES 137–146 )................................................ 28. Promecidia Lelej, 1996

25. Mesosoma broadest in pronotum or pronotum and propodeum with equal width ( Figs 21, 23, 24 View FIGURES 1–28 )..................... 26

- Mesosoma definitely broadest in propodeum ( Figs 20, 26, 27 View FIGURES 1–28 )................................................. 30

26. Mesosoma black; metasomal segment 2 except posterior border, reddish................ 29. Taiwanomyrme Tsuneki, 1993

- Mesosoma ferruginous-red; metasomal segment 2 black...................................................... 27

27. Clypeus anteriorly strongly emarginate and bidentate.................................... 26. Pagdenidia Lelej, 1996

- Clypeus anteriorly at most weakly emarginate, with two weak denticles.......................................... 28

28. Scutellar scale broad, 1/3–1/4 × propodeal width; pygidial area finely rugoso-punctate.......... 32. Zavatilla Tsuneki, 1993

- Scutellar scale narrow; pygidial area mostly smooth, at most micropunctate...................................... 29

29. T2 with wide apical pale pale setal band ( Fig. 21 View FIGURES 1–28 )..................................... 2 3. Krombeinidia Lelej, 1996

- T2 without apical pale setal band, at most with sparse pale fringe ( Fig. 24 View FIGURES 1–28 )....................................................................................................... 27. Petersenidia Lelej in Lelej & Yamane, 1992

30. Pygidial area irregularly rugose throughout ( Fig. 124 View FIGURES 115–124 )........................... 31. Wallacidia Lelej & Brothers, 2008

- Pygidial area variable, if mostly rugose, then having apex smooth or microreticulate ( Figs 117, 121 View FIGURES 115–124 , 151, 154 View FIGURES 147–154 )........... 31

31. Pygidial area longitudinally striate ( Figs 154 View FIGURES 147–154 , 164 View FIGURES 155–164 ), longitudinally rugose basally ( Fig. 160 View FIGURES 155–164 ), or rugose in basal 0.75 ( Figs 151 View FIGURES 147–154 , 157 View FIGURES 155–164 )....................................................................... 30. Trogaspidia Ashmead, 1899

- Pygidial area irregularly rugose in basal half and smooth in apical half ( Fig. 121 View FIGURES 115–124 ) or entirely microreticulate ( Fig. 117 View FIGURES 115–124 )................................................................................... 22. Eotrogaspidia Lelej, 1996