Acylophorus madecassus, Janák, 2018

Janák, Jiří, 2018, Revision of the genus Acylophorus Nordmann, 1837 from Madagascar and Mascarene Islands (Coleoptera: Staphylinidae: Staphylininae: Acylophorina), Zootaxa 4462 (2), pp. 151-191: 167

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Acylophorus madecassus

sp. nov.

Acylophorus madecassus   sp. nov.

( Figs. 67–77 View FIGURES 67–73 View FIGURES 74–77 , 175 View FIGURES 172–177 )

Type locality. Madagascar, Sambava , near Anovona river, cca 14 15' 54" S 50 09' 02" E.  

Type material. Holotype ♂: MADAGASCAR: Madagascar Est , Sambava , Riv. [Rivière = river] Anovona env., 0–20 m, 3.– 4.3.1996, rizières/bord de la forêt [rice fields/forest edge], J. Janák & P. Moravec lgt. ( JJRC)   .

Description. Body length 7.5 mm, forebody length 3.4 mm. Body black ( Fig. 67 View FIGURES 67–73 ), pronotum and abdomen brown, abdominal tergites markedly iridescent; antennae brown with base of first two segments and segments 7–11 reddish yellow; legs reddish brown, tarsi and maxillar palpi reddish yellow, abdomen gold iridescent.

Head subrounded ( Fig. 68 View FIGURES 67–73 ), about as long as wide (L/W=1.01), markedly widened behind eyes, with two pairs of interocular setae. Temples longer than eyes (R=1,56). Dorsal side of head shining, without scatered extremely fine micropunctures. Dense pale grey pubescece behind eyes. Four postocular setae visible on each side, additional seta on hind margin of eye present. Left mandible with two teeth, right mandible with three teeth ( Fig. 69 View FIGURES 67–73 ). Maxillary palpi with terminal segment long and relativelly narrow, pubescent, asymmetric, markedly wider and longer than glabrous penultimate segment which is elongate ( Fig. 70 View FIGURES 67–73 ). First segment of antenna nearly as long as next four; segments 1 to 4 elongate, segment 5 markedly longer than wide (R=1.61), segment 8 about as long as wide, segment 9 markedly transverse, segment 10 markedly transverse (R=0.62).

Pronotum slightly transverse (R=1.14), markedly widened behind with sides rounded, widest in basal half ( Fig. 68 View FIGURES 67–73 ), shining, with scattered extremely fine micropunctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra slightly transverse (R=1.36) with short, pale, but not shining pubescence arising from dense aspirate punctures. Apical fringe of bristles markedly longer than pubescence on other parts of the elytra. Abdominal tergites with moderately long and moderately sparse pubescense, arising from aspirate punctures that are irregularly distributed across each tergite and more coarse but markedly sparser than on elytra.

Meso- and metatarsi with empodial setae longer than claws (similar as in Figs. 9, 10 View FIGURES 9–17 ).

Male. Sternite 9 very long and narrow ( Fig. 77 View FIGURES 74–77 ), apex of tergite 10 narrowly rounded, with very long setae ( Fig. 76 View FIGURES 74–77 ). Aedeagus ( Figs. 71–75 View FIGURES 67–73 View FIGURES 74–77 ) 0.92 mm long. Parameres bilobed, lobes widely separated, narrowed apically, sensory pegs confusedly arranged in apical half of both lobes ( Fig. 75 View FIGURES 74–77 ). Median lobe of aedeagus markedly longer than paramere, apex broadly rounded ( Figs. 73 View FIGURES 67–73 , 75 View FIGURES 74–77 ).

Female. Unknown.

Differential diagnosis. Acylophorus madegassus   sp. nov. differs from all hirtherto known Aftrotropical Acylophorus   by the very long temples and by the different shape of aedeagus.