Harmothoe bellani, Barnich & Fiege, 2000

Barnich, R. & Fiege, D., 2000, Revision of the Mediterranean species of Harmothoe Kinberg, 1856 Lagisca Malmgren, 1865 (Polychaeta: Polynoidae: Polynoinae) descriptions of a new genus and a new species, Journal of Natural History 34, pp. 1889-1938 : 1920-1921

Harmothoe bellani sp. nov.

(fi gure 16A -D)

Harmothoe reticulata: Fauvel, 1913: 270 [part].

Harmothoe goreensis: Amoureux, Rullier and Fishelson, 1978: 64 .

Type material. Harmothoe bellani , HOLOTYPE: 10 September 1997, Banyuls, posidonies, preÁs du Labo, 10 m, leg. D. Fiege and R. Barnich, SMF 9087 . PARATYPE:

spm., 21 June 1994, Banyuls, Hafen, 2-5 m, leg. H. Zibrowius, SMF 9088 .

Additional material. 2 spms., 21.6.94, Banyuls Hafen, 2-5 m, leg. Zibrowius, SMF 9089 (in bad shape). 1 spm., 1910, St. 1260, Monaco, det. Fauvel as H. reticulata in Fauvel 1913), IEA I 25’ (slide). 1 spm., 1973, Mer Rouge, Marsa-el-At., golfe d’Elat, 25 m dans une eÂponge, in Amoureux et al. (1978), det. as H. goreensis, IEA

99 (slide).

Diagnosis. Anterior pair of eyes dorsolaterally on widest part of prostomium. Elytra: margin with long fringing papillae; surface covered by thorn-shaped pointed

Revision of the Mediterranean Harmothoe and L agisca 1921 IG. 16. Harmothoe bellani sp. nov. (holotype, SMF 9087). (A) Anterior end; style of median antenna missing, right dorsal tentacular cirrus regenerating; dorsal view; (B) right anterior elytron; (C) detail of posterior margin of same; (D) style of dorsal cirrus (scales 5 1 mm for A, B; 100 M m for C; 500 M m for D).

microtubercles and few scattered papillae. Dorsal and ventral tentacular cirri and dorsal cirri papillate, tapering.

Description. Prostomium bilobed, with distinct cephalic peaks; median antenna with large ceratophore in anterior notch, style papillate, tapering, pigmented basally and with a pigmented band distally; lateral antennae with small ceratophores inserted ventrally and with short, tapering papillate styles, each pigmented basally and with

pigmented band distally; palps papillate, about as long as median antenna and tentacular cirri; anterior pair of eyes dorsolaterally on widest part of prostomium, posterior pair dorsally near hind margin. Tentaculophores inserted laterally to prostomium, each with two chaetae and a pair of papillate, tapering dorsal and ventral tentacular cirri, each pigmented basally and with two pigmented bands distally. Second segment with fi rst pair of elytra, biramous parapodia, and long ventral buccal cirri, similar to tentacular cirri (fi gure 16A).

Fifteen pairs of elytra; elytral margin with long fringing papillae; surface covered

thorn-shaped pointed microtubercles and few scattered papillae; pigmentation: mottled, brownish in alcohol (fi gure 16B, C). Styles of dorsal cirri papillate, tapering, extending beyond neurochaetae, each pigmented basally and with two pigmented bands distally (fi gure 16D); styles of ventral cirri papillate, tapering, shorter than neuropodia.

Parapodia biramous, both rami with single aciculum penetrating epidermis; notopodia with short, rounded presetal and longer, pointed postsetal acicular lobe, neuropodia with subconical presetal acicular lobe with digitiform supra-acicular process and shorter rounded postsetal lobe. Notochaetae stouter than neurochaetae, with distinct rows of spines and blunt tips; neurochaetae with distinct rows of spines 1922 only in distal part, upper bidentate with small, stout secondary tooth distally, lower