Eremotylus, Randall T. Schuh & Michael D. Schwartz, 2016

Randall T. Schuh & Michael D. Schwartz, 2016, Nineteen new genera and 82 new species of Cremnorrhinina from Australia, including analyses of host relationships and distributions (Insecta: Hemiptera: Miridae: Phylininae: Cremnorrhinini), Bulletin of the American Museum of Natural History 401, pp. 1-279 : 77-79

Eremotylus , new genus

TYPE SPECIES: Eremotylus stuarti , new species.

DIAGNOSIS: Recognized by the ovoid to elongate-ovoid body form and green to yellowish coloration; dorsum sometimes with a dark spot on membrane at apex of cells or on corium at inner angle of cuneus (pl. 8); endosoma sigmoid, usually with one short and one long spine apically, sometimes with broad subapical flap (figs. 26–30, pl. 9). Possibly confused with Gyrophallus on form of apical endosomal spines, but that taxon always with endosoma twisted (like a rope) (pls. 13, 15) as opposed to being just sigmoid. Gyrophallus also always with conspicuous apical endosomal membranes with some denticles, attributes never seen in Eremotylus .

(pl. 8): Green to yellowish, including all appendages, hemelytron sometimes nearly transparent; hemelytron either without markings, or occasionally corium with infuscate mark adjacent to inner angle of cuneus and/ or membrane with infuscate spot at apex of cells. SURFACE AND VESTITURE (pl. 8): Dorsum smooth, polished and weakly to strongly shining. Dorsum clothed with recumbent simple setae, sometimes pale, but often dark. STRUCTURE: Head (pl. 8): Short, transverse; frons at most weakly surpassing anterior margin of eyes; eyes moderately large, usually weakly bulging, confluent with anterior margin of pronotum; occupying about two-thirds height of head; antenna inserted near ventral margin of eye; antennal segment 2 ranging from short to long. Thorax (pl. 8): Pronotum with lateral margins nearly straight to weakly sinuous, anterior lobe short, calli weakly to distinctly demarcated along posterior margin, posterior lobe at most weakly elevated, posterior margin straight to weakly concave, with rounded humeral angles; mesoscutum weakly to broadly exposed; scutellum triangular, flat. Hemelytron: Short to moderately elongate, corial margin nearly straight to moderately convex. GENITALIA (figs. 26–30, pl. 9): Pygophore: Broadly or elongate conical, dorsal surface anteriad of aperture sometimes with narrow tubercle on left side, rarely also with bump on right side; sometimes with conspicuous long bristle bilaterally near paramere insertions. Endosoma: Sigmoid, apical portion sometimes bent to left in dorsal view; ventral strap usually terminating proximad of secondary gonopore and dorsal strap usually bifid proximal to or distad of secondary gonopore with one long and one short apical spine; secondary gonopore well sclerotized, circular, situated subapically. Phallotheca: Apical portion narrow, dorsal margin with narrow crest or without crest, aperture usually narrow, slitlike, sometimes wider, situated on anterodorsal margin and continuous across apex, or on posterior side; basal portion reaching to middle of pygophore in situ. Parameres: Left paramere usually relatively elongate in dorsal view, rarely more compact; usually with medial portion of dorsoposterior margin undulating, strongly elevated above posterior and anterior processes, sometimes dorsoposterior margin practically straight; posterior process usually with bent distal one-half and undulating lateral margin in dorsal view, sometimes posterior process straight; anterior process variable. Right paramere usually of moderate length, usually with one rather broad truncate apical projection.

FEMALE (pl. 8): Total length 2.64–3.81, pronotum width 0.81–1.09. Coloration and structure as in male, except vertex wider, antennal segment 2 usually more slender, and body form more strongly ovoid. GENI- TALIA (pl. 44): Subgenital plate of sternite 6: Concave medially. Vestibular sclerites: Moderately large, attaining anterior edge of dorsal labiate plate. First gonapophyses: Moderate size, wedge-shaped basal blocks. Ventral labiate plate: Platelike, medial anteroventral extension narrow, covering anterior surface of basal structures. Dorsal labiate plate: Relatively short longitudinally. Sclerotized rings: Moderately large, relatively flat, thick walled, teardrop shaped, medial angle strongly attenuate. Posteromedial region: Surface without apparent microstructure. Anterolateral region: Anterior margin even with anterior edge of sclerotized rings. Posterior wall: Intersegmental structure: Not differentiated from connecting membrane. Interramal sclerites: Weakly sclerotized, lateral sclerites narrow, medial sclerite apparently absent.

ETYMOLOGY: From the Greek, eremos, “solitary or lonely,” and tylus, “knob, nose,” in reference to the projecting face of most members of the Cremnorrhinina; masculine.