Cyrnellus fraternus ( Banks 1905 )

Rueda Martín, Paola A., Gibon, Francois-Marie & Sganga, Julieta V., 2021, New species and records of the Neotropical genus Cyrnellus (Trichoptera: Polycentropodidae) and the re-establishment of C. minimus and C. marginalis, Zootaxa 5082 (1), pp. 15-29 : 20

Cyrnellus fraternus ( Banks 1905)

Figs 3E–3K

Nyctiophylax fraternus Banks 1905: 17 (original description); Ross 1938: 12 (designation of female lectotype).

Cyrnellus fraternus ; Flint 1964: 469–471 (synonymy, description of larva); Holzenthal & Calor 2017: 419 (catalogue, synonymy).

Discussion. This species was described originally based on a female from Plummer’s Island, Maryland, USA. The females of C. fraternus and C. marginalis are currently indistinguishable. Flint (1964) thought that C. fraternus and C. marginalis are synonyms and that C. fraternus is the only species of Cyrnellus in North America. That “single species” was thought to be widely distributed in North America, occurring in at least Alabama, Arkansas, Illinois, Kentucky, Michigan, Minnesota, Missouri, Ohio, Oklahoma, Tennessee, and Wisconsin ( Ross 1944). Records from these and many additional states have been reported since 1944 (Rasmussen & Morse 2021). Based on our conclusion that males of the two species are distinguishable (see diagnosis of C. marginalis , Figs 3E–3K ), identifications of voucher specimens from those many reports will need to be reviewed in order to determine the ranges of these two species in North America.

Figures

FIGURE 3. Cyrnellus spp., male genitalia. 3A–3D, Cyrnellus marginalis Banks 1930: 3A, left lateral; 3B, right inferior appendage, ventral; 3C, phallus, left lateral; 3D, phallus, dorsal. 3E–3K, Cyrnellus fraternus (Banks 1905): 3E, left lateral; 3F, right inferior appendage, ventral; 3G, phallus, left lateral; 3H, phallus, dorsal; 3I–3K, variations of the right inferior appendage, ventral. ap = apex of an inferior appendage; as = subapicomesal spine; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process.