Cagiva Pogue, 2019

Brown, John W., 2019, New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae), Zootaxa 4671 (2), pp. 195-222 : 201

Cagiva Pogue , new genus

Type species: Phalonia cephalanthana Heinrich, 1921 , by monotypy.

Cagiva Pogue and Mickevich, 1990: 322 (nomen nudum).

Cochylini New Genus 2 [ cephalanthana group]: Brown 2005: 208; Metzler and Brown 2014: 278.

Cagiva is proposed for Phalonia cephalanthana , a species that has long defied confident generic assignment. The species was described in Phalonia by Heinrich (1921) and placed in “incertae sedis” by Powell (1983). The monotypic genus Cagiva was proposed by Pogue (1986) in his unpublished Ph.D. dissertation. Brown (2005) and Metzler and Brown (2014) followed that concept, designating the manuscript name Cagiva as “ Cochylini New Genus”.

Diagnosis. Putative autapomorphies for Cagiva include the prominent, well-sclerotized socii, and the distally bifurcate phallus. The genus is further characterized by a unique combination of characters, some of which are shared with other genera: the 3-segmented maxillary palpi (shared with Thyraylia , Nycthia , and few others), the absence of cornuti in the vesica (shared with Eupinivora Brown, 2013 , Honca , and Nycthia ), and the connate condition of hindwing veins M 3 and CuA 1 (shared with a few other genera). As in most Cochylina, the male of Cagiva lacks a forewing costal fold and has a well-developed hindwing costal roll.

Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna 0.8–1.0 times flagellomere diameter in male, much shorter in female; labial palpus porrect, combined length of all segments 1.1–1.2 times diameter of compound eye, segment III exposed; maxillary palpus 3-segmented. Thorax: Posterior crest present; lateral scale tufts of metanotum flat. Forewing ( Fig. 2 ) length 4.0– 4.7 mm, length 2.4–2.8 times width, slightly wider in female; costa gently arched from base to apex, without costal fold in male, apex abruptly rounded; termen straight; Sc slightly less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 originating nearer R 3 than R 1; R 5 ending at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.65 length of discal cell; CuP absent; A1+2 stalked at ca. 0.5 length. Hindwing moderately broad, length 2.3–2.8 times width; costa straight, with costal roll extending about 0.65 length of costa in male; apex rounded; termen slightly concave below apex; Sc+R 1 ca. 0.25 wing length in male, 0.65–0.75 wing length in female; Rs and M 1 stalked ca. 0.7 length of M 1; M 3 and CuA 1 connate; CuA 2 originating ca. 0.65 length of discal cell; frenulum with one acanthus in male, two in female.Abdomen: Unmodified. Male genitalia ( Fig. 21 ) with uncus absent; socius prominent, well sclerotized, setose, distally elongate and curved mesially; transtilla well developed, median process a small rounded tooth; valva length 1.4‒1.5 times width; costa sclerotized, relatively straight; sacculus well defined, confined to basal 0.33 of venter of valva, terminating in a rounded lobe; vinculum arms weakly join distally by thin membrane. Phallus large, ca. 1.3 times length of valva, straight, parallel-sided in basal 0.5, broadened in distal 0.5, bifurcate apically, asymmetric; vesica without cornuti. Female genitalia ( Fig. 35 ) with telescopic ovipositor; papillae anales narrow, weakly sclerotized, setose, joined posteriorly; length of apophyses anteriores ca. 2.0 times that of apophyses posteriores, ventral branch of apophyses anteriores reduced, apophyses with rounded swelling at anterior tip; sterigma complicated, comprised of weakly sclerotized sacs, some bearing setae, and a crescent-shaped plate posterad of ostium; sterigma not attached to apophyses anteriores; ductus bursae relatively broad, rather uniform in width, ca. 0.65 as long as corpus bursae, differentiated from corpus bursae, with caudal 0.7 weakly sclerotized; accessory bursa originating near ostium; corpus bursae oblong, covered with minute spicules, signum lacking.

Etymology. The name is an arbitrary combination of letters and is considered feminine in gender.

Figures

FIGURES 1–10. Adults of Cochylini genera. 1. Anacochylidia maderana, Madera Canyon, Santa Rita Mountains, Arizona (EME). 2. Cagiva cephalanthana, Dorman Lake, Oktibbeha Co., Mississippi (USNM). 3. Atroposia oenotherana, Houston, Harris Co., Texas (USNM). 4. Atroposia oenotherana, Radisson, Onondaga Co., New York (USNM). 5. Cybilla hubbardana, Shafter, Presidio Co., Texas (USNM). 6. Cybilla hubbardana, Lordsburg, Hidalgo Co., New Mexico (CNC). 7. Honca grandis, Chimney Gulch, Golden, Colorado (USNM). 8. Pogospinia floridana, Long Key State Park, Monroe Co., Florida (USNM). 9. Nycthia pimama, Portal, Cochise Co., Arizona (USNM). 10. Nycthia yuccatana, Falcon State Park, Starr Co., Texas (USNM).

FIGURES 19–26. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249.

FIGURES 33–38. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660.