Heterocerus intermuralis, (Pacheco)

King, Jonas G. & Lago, Paul K., 2012, The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States, Insecta Mundi 2012 (275), pp. 1-53 : 9-10

9(5). Elytra grayish-brown; pronotum darker brown, often with pale margins ( Fig. 12 ); anterior end of phallobase about half the maximum width of the phallobase ( Fig. 34 ) ................................... ....................................................................................................... 7. H. intermuralis (Pacheco) —

Elytra testaceous to dark brown; pronotum usually darker than elytra, coloration of margins variable; anterior end of phallobase never constricted to less than 80% of maximum width .... ......................................................................................................................................................... 10 10(9).

— Parameres quadrate ( Fig. 33 ); testaceous dorsally; hypermandibulate males unknown. Uncommon, known only from southern Texas and southwestern Louisiana .......... EL H. glicki Pacheco

Parameres triangular; dorsal color variable; hypermandibulate males relatively common. Widespread in the United States.................................................................................................. 11

Figures

Figures 11-17. Habiti of large Heterocerus species lacking post-mesocoxal lines. 11) H. pallidus (male is hypermandibulate). 12) H.intermuralis (male is hypermandibulate). 13) H.fatuus. 14) H. gnatho. 15) H.longilobulus. 16) H. sandersoni. 17) H. angustatus.Scale line = 1mm.

Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus.