Tipula (Vestiplex) tchukchi Alexander

Starkevich, Pavel, Podenas, Sigitas & Gelhaus, Jon K., 2020, Taxonomic review of Tipula (Vestiplex Bezzi) crane flies (Diptera: Tipulidae) in Mongolia, Zootaxa 4837 (1), pp. 1-88 : 55-58

publication ID	https://doi.org/ 10.11646/zootaxa.4837.1.1
publication LSID	lsid:zoobank.org:pub:001B5DF8-A24D-4353-8BEA-8B181BE9D279
DOI	https://doi.org/10.5281/zenodo.4454933
persistent identifier	https://treatment.plazi.org/id/03E187A2-9310-A850-6FEB-FF6F212FC30A
treatment provided by	Plazi
scientific name	Tipula (Vestiplex) tchukchi Alexander
status

Treatment

( Figs 20 View FIGURES 14–27 , 117–128 View FIGURES 117–123 View FIGURES 125–128 ; Map 9 View MAPS 7–9 )

Tipula (Vestiplex) tchukchi Alexander, 1934b: 408; 1935: 118.

Tipula (Vestiplex) laccata: Lackschewitz, 1936: 264 (partim); Mannheims, 1953: 132 (partim); Savchenko, 1964: 204 (partim); nec Lundström & Frey, 1916 (after Mannheims 1971).

Tipula (Vestiplex) bo Mannheims, 1967: 148 ; Mannheims & Savchenko, 1973: 172 (synonymy).

Tipula (Vestiplex) laccata obtusidens Savchenko, 1964: 205 .

Tipula (Vestiplex) thukchi obtusidens: Starkevich & Paramonov, 2016: 81 ; Starkevich & Paramonov, 2016: 81 (synonymy).

Type material examined. Tipula (Vestiplex) tchukchi Alexander : HOLOTYPE ♂: RUSSIA, Chukchi Peninsula, Chukotka Autonomous Okrug, Markovo township near Anadyr town , 6.vii.1896, Gondatti ( ZIN) . PARATYPES: 1 ♀, topotypic ( ZIN) ; 1 ♂, Kamchatka Kray, mouth of Kichiga River, 27.vi.1910, Skorikov ( ZIN) .

Additional material examined. MONGOLIA. Bulgan Aimag: 1 ♀, between Somon Chischig-Öndör and Somon Orchon, 23 km NNE von Somon Chischig-Öndör, 1390 m, Exp. Dr. Z. Kaszab , 1968, Nr. 965, 15.vi.– 23.vii.1968 ( HNHM) . Hovsgol Aimag: 1 ♂, NE shore of Hovsgol Lake, Noyon valley , riparian zone, N5°12′05.5″, E100°48′26.2″, 1709 m, 8.vii.2002, O. Yadamsuren GoogleMaps . Ovorkhangai Aimag: 5 ♂, Bat-Olziy Soum, Ovtiyn Gol, 10 km SE Bat-Olziy, N46.75972, E102.30905, 1847 m, 8.vii.2011, S. Podenas, MAIS2011070803 GoogleMaps . Selenge Aimag: 1 ♀, Shaamar Soum, Orkhon River at Delgerkhaan Bridge , N50.06476, E106.13388, 609 m, 27.vii.2003, SRPT, SRP03072701 GoogleMaps . Tov Aimag: 1 ♂, Erdene Soum, Baruum Bayan Gol (river), 48°06′N, 107°45′E, 29.vi.1998, Sh. Enkhetsetseg ; 1 ♂, 1 ♀, Erdene Soum, Gorkhi Terelj NP, Khokh Chuluutiin Gol , N48.12675, E107.57996, 1680 m, 6.vii.2003, SRPT, SRP03070604 GoogleMaps ; 4 ♂, 5 ♀, Erdene Soum, Gorkhi Terelj NP, unnamed trib. of Tuul River on West side, 1.6 km upstream from Daichin crossing, N48.21780, E107.90392, 1594 m, 9.vii.2003, SRPT, SRP03070902 GoogleMaps ; 1 ♀, Erdene Soum, Gorkhi Terelj NP, mouth of Khag River at confluence with Tuul River , N48.25861, E107.90251, 1608 m, 10.vii.2003, SRPT, SRP03071002 GoogleMaps ; 1 ♂, Erdene Soum, Gorkhi Terelj NP, stream 0.5 km N Gorkhi Davaa ( Pass ), N47.94981, E107.45511, 1593 m, 11.vii.2011, S. Podenas, MAIS2011071101 GoogleMaps ; 1 ♂, Erdene Soum, Shar Bulag, Yellow Spring , 26 km W Mongonmorit, N48.14814, E108.07956, 1629 m, 13.vii.2011, S. Podenas, MAIS2011071301 GoogleMaps .

Elevation range in Mongolia. Adults were collected at altitudes ranging from 600 m to 1850 m.

Period of activity. Adults are active from the end of June through to the end of July.

Known distribution. Finland, Mongolia ( Map 9 View MAPS 7–9 ), Russia (eastern part of Siberia, Far East) and Sweden.

Redescription. Male. Body length 13.5–15.8 mm, wing length 15.0–16.0 mm. General body coloration blackish yellow.

Head. Gray, vertex and occiput gray with brown median line. Coloration of rostrum varies from yellow to brown, dorsally dusted with gray. Nasus short. Antenna 13-segmented, if bent backward extending beyond wing base. Scape, pedicel and flagellomere 1 yellowish, succeeding flagellomeres brown. Each flagellomere except first one with basal enlargement and moderately incised. Apical flagellomere small, reduced. Verticils shorter than corresponding flagellomeres. Palpus dark brown.

Thorax. Blackish gray pruinose. Pronotum blackish, gray dusted, with brown median line. Prescutum and presutural scutum black, grey pruinose with 4 longitudinal stripes bordered by brown. Intermediate pair fused into brown median line. Interspace between median and lateral stripes light gray with light setae. Postsutural scutum blackish, gray pruinose with median line. Scutal lobe with 2 spots bordered by brown. Scutellum brown with median line. Postnotum grey dusted with midline and long setae. Pleura blackish, grey pruinose. Coxae blackish, grey pruinose. Trochanters, femora and tibiae yellowish. Tarsal segments brown. Distal part of femora and tibiae dark brown. Tarsal claws toothed. Wing slightly patterned with brown. Halter with yellowish stem and brown knob.

Abdomen. Brownish yellow. Tergites with lateral margins narrowly pale. Tergite 1 dark brown, dusted with gray. Dorsal stripe on tergites 3–5 broadly interrupted. Tergite 6 black in posterior half, tergites 7–8 black. Sternite 1 brown, sparsely dusted with gray. Lateral stripe distinct. Sternites 2–5 and sternite 6 basally at about one-third length brownish yellow, succeeding sternites black.

Hypopygium. Black. Tergite 9 forming large concave sclerotised saucer roughly rectangular in outline ( Fig. 117 View FIGURES 117–123 ). Main body of tergal saucer brown with blackened rim. Posterior margin of tergal saucer broadly emarginated, with median U-shaped notch, 2 yellow rounded projections and black median transverse sclerotisation, small denticles on either side. Lateral angle of tergal saucer obtuse, broadly truncated. Anterior and lateral portions of tergal saucer raised into sclerotised border with obtuse anterior angles. Border laterally produced into obtuse point directed caudad and situated under lateral angle of tergal saucer. Gonocoxite unarmed ( Fig. 118 View FIGURES 117–123 ). Outer gonostylus flattened, slightly curved with apex rounded ( Fig. 119 View FIGURES 117–123 ). Inner gonostylus in shape of curved plate, bidentate at apex, with obtuse tooth dorsally, beak claw-shaped ( Fig. 120 View FIGURES 117–123 ). Gonocoxal fragment large with lateral and medial sclerites well-developed ( Fig. 121 View FIGURES 117–123 ). Medial sclerites fused, anterior apodeme narrowed, posterior part broadened and arched, posterior apodeme with apex rounded and medially incised. Lateral sclerite large and bilobed, expanded at base. Aedeagal guide relatively narrow tube-shaped ( Fig. 122 View FIGURES 117–123 ). Sperm pump with central vesicle small and flattened ( Fig. 123 View FIGURES 117–123 ). Compressor apodeme with median incision. Posterior immovable apodeme longer than compressor apodeme, relatively narrow. Anterior immovable apodeme rounded. Aedeagus shaped as moderately long tube, about 3.8 times as long as sperm pump, basally dark brown, medially brown passing into yellow towards apex. Distal part ventrally membranous, shovel-shaped (Fig. 124).

Female. Body length 17.3–24.5 mm, wing length 12.5–14.5 mm. Antenna short, if bent backward extending beyond pronotum. Antenna yellowish, flagellomeres cylindrical. Abdominal stripe broad, distinct.

Ovipositor ( Figs 125–128 View FIGURES 125–128 ). Tergite 10 shiny, brownish-yellow. Cercus brownish-yellow, slightly shorter than tergite 10, curved dorsally with apical incision, outer margin with rough and distinct serration ( Fig. 125 View FIGURES 125–128 ). Hypovalva in shape of yellow flattened filament with broadened base and short trichia at tip ( Fig. 126 View FIGURES 125–128 ). Median incision between hypovalvae deeper than posterior margin of sternite 8. Lateral incision relatively deep, about twice as deep as maximum width. Lateral angle of sternite 8 obtuse. Sternite 9 with anterior part broad, inner edge with submedian triangular extension, posterior part straight ( Fig. 127 View FIGURES 125–128 ). Furca anteriorly narrowed, posteriorly broad ( Fig. 128 View FIGURES 125–128 ). Bursa copulatrix with spermathecal ducts sclerotised at base, in shape of nearly straight dark brown process ( Fig. 128 View FIGURES 125–128 ). Wall of bursa copulatrix on connection site with spermathecal ducts sclerotised. Sclerotisation of all 3 spermathecal ducts connected and forming complete dark brown ring. Cul-de-sac part of bursa copulatrix roughly straight. Spermatheca pear-shaped ( Fig. 26 View FIGURES 14–27 ).