Tipula (Vestiplex) longitudinalis Nielsen

Starkevich, Pavel, Podenas, Sigitas & Gelhaus, Jon K., 2020, Taxonomic review of Tipula (Vestiplex Bezzi) crane flies (Diptera: Tipulidae) in Mongolia, Zootaxa 4837 (1), pp. 1-88 : 69-76

Tipula (Vestiplex) longitudinalis Nielsen

( Figs 3 , 4 , 15 , 163–173 ; Map 13 )

Tipula longitudinalis Nielsen, 1929: 49 .

Tipula (Vestiplex) longitudinalis: Savchenko, 1960: 196 ; 1964: 213; Oosterbroek & Theowald, 1992: 156.

Material examined. MONGOLIA. Arkhangay Aimag: 3 ♂, Khotont Soum, trib. of Tsagaan Sumiin/Jarantain Gol, ~ 43 km SW Khotont, N47.07888, E102.16608, 1699 m, 9–10.vii.2004, SRPT, SRP 04070902; 28 ♂, Tsenkher Soum, Tsetseleg Gol ~ 17 km SW Tavanbulag, N47.26999, E101.80227, 1684 m, 10–12.vii.2004, SRPT, SRP 04071002; 1 ♂, Tsenkher soum, Uliin Gol in Budent valley ~ 60 km SSW Tavanbulag, N46.97927, E101.69877, 2075 m, 11.vii.2004, SRPT, SRP 04071101; 5 ♂, Tsenkher Soum, Tsetserleg Gol ~ 50 km SSW Tavanbulag, N47.04409, E101.76352, 1902 m, 11.vii.2004, SRPT, SRP 04071102; 22 ♂, Ikhtamir Soum, NW Khoit Tamir Gol ~ 25 km SW Ikhtamir, N47.50523, E100.93208, 1728 m, 14–15.vii.2004, SRPT, SRP 04071402; 10 ♂, Chuluut Soum, Khurmen/Davaat Gol 15 km SSE Chuluut/Jargalant, N47.42580, E100.30130, 2104 m, 15–16.vii.2004, SRPT, SRP 04071503; 10 ♂, Ikhtamir Soum, Khoit Tamir Gol ~ 29 km SW Ikhtamir, N47.48567, E100.87875, 1749 m, 15.vii.2004, SRPT, SRP 04071501; 3 ♂, Chuluut Soum, Khanui Gol ~ 60 km SW Ikhtamir, N47.50949, E100.57510, 1981 m, 15.vii.2004, SRPT, SRP 04071502; 3 ♂, Chuluut Soum, Tsohiotiin Davaa (mountain pass), N47.68999, E100.21012, 2256 m, 17.vii.2004, J.K. Gelhaus #969; 1 ♂, Chuluut Soum, Chuluutin Gol 40 km N Chuluut/Jargalant, N47.890440, E100.31810, 1937 m, 17.vii.2004, SRPT, SRP 04071701; 7 ♂, Tariat Soum, Nariin Gichgenii Gol/Urd Terkh Gol ~ 5 km SE Khorgo/Tariat, N48.12661, E99.94128, 2019 m, 17–18.vii.2004, SRPT, SRP 04071702; 2 ♂, Tariat Soum, Terkhiin Tsagan Nuur ~ 8 km W Khorgo/Tariat, N48.14404, E99.78300, 2057 m, 18.vii.2004, SRPT, SRP 04071801; 3 ♂, 1 ♀, Khangay Soum, Khoit Ekhnii Gol ~ 13 km SW Khunt, N47.75747, E99.35827, 2215 m, 18–19.vii.2004, SRPT, SRP 04071803; 1 ♂, Khashaant Soum, Tarnay Gol, Mongol Els, 24 km W Rashaant, N47.35471, E103.62062, 1234 m, 1.vii.2011, C. R. Nelson & T.A. McKnight & MAIST, MAIS 2011070102. Bayan Olgii Aimag: 1 ♀, Ulaanhus Soum, Ulastai Gol 2 km E Khokh-Ereg Military Outpost, N49.31867, E88.36612, 2392 m, 8.vii.2008, R.W. Bouchard, Jr., MAIS 08070802; 1 ♂, Tsengel Soum, Sul Uuliin Gol, trib. of Tsagaan Gol, Altai-Tavan Bogd NP, N49.10448, E88.04845, 2796 m, 10.vii.2008, S. Podenas, MAIS 08071001; 9 ♂, Tsengel Soum, Tsagaan Gol, 20 km W Zagast Nuur Bag, Entrance to Altai-Tavan Bogd NP, N49.09161, E88.10436, 2410 m, 9–12.vii.2008, S. Podenas, MAIS 08070902; 7 ♂, 2 ♀, Tsengel Soum, Urt Khuiten Gol 6 km SW Asgut Uul, N48.72627, E88.40073, 2427 m, 14.vii.2008, R.W. Bouchard, Jr. MAIS 08071401; 4 ♂, 1 ♀, Tsengel Soum, Khoton Nuur 15 km NW Syrgal, N48.66722, E88.29927, 2086 m, 14–15.vii.2008, S. Podenas, MAIS 08071402; 8 ♂, Tsengel Soum, Rashaanii Gol, at bridge, trib. of Tsagaan Us Gol, N48.75565, E88.15266, 2098 m, 15.vii.2008, S. Podenas, MAIS 08071501; 4 ♂, Tsengel Soum, Tsagaan Us Gol, 0.5 km upstream of confluence with Rhashaanii Gol, N48.75321, E88.14850, 2090 m, 15.vii.2008, MAIST, MAIS 08071502; 4 ♂, 1 ♀, Tsengel Soum, Ikh Turgenii Gol, 6 km SSW Syrgal, N48.54395, E88.41477, 2140 m, 15–16.vii.2008, MAIST, MAIS 08071503; 55 ♂, 1 ♀, Tsengel Soum, Khar Ovoo Gol, draining Khelkhee Lake, N48.62918, E88.28554, 2147 m, 15–16.vii.2008, MAIS 2008071601; 14 ♂, Tsengel Soum, Partizanii Bulag below Khazabai Sala (waterfall), 2148 m, N48.61734, E88.31315, 16.vii.2008, R.W. Bouchard. Jr., MAIS 08071602; 4 ♂, Tsengel Soum, Sumdairag Gol, 12 km S Syrgai, 2129 m, N48.50679, E88.50977, 16.vii.2008, J. Morse, Suvdaa, S. Podenas, MAIS 08071604; 1 ♂, Tsengel Soum, Syrgali Gol, bridge over outflow between Khoton Nuur and Khurgen Nuur at Syrgal, 2095 m, 16.vii.2008, Suvdaa, MAIS 08071603; 16 ♂, Sagsay Soum, Yamaatiin Gol, 17 km SW Dayan Military Base, N48.16728, E88.85435, 2073 m, 17.vii.2008, MAIS 2008071703; 2 ♂, Ulaanhus Soum, Odgoi Gol, 21 km SE Syrgali, N48.44036, E88.60204, 2443 m, 17.vii.2008, MAIST, MAIS 08071701; 4 ♂, 1 ♀, Sagsay Soum, Ikh Khanajashiin Gol, 17 km by road N Dayan, N48.39223, E88.88369, 2184 m, 18.vii.2008, MAIST, MAIS 08071802. Bayankhongor Aimag: 2 ♂, Erdenesogt Soum, Erkhetkhayrkhan Gol, crossing to Khokh Gol, N46.62626, E101.36736, 2500 m, 5.vii.2011, S. Podenas, MAIS 2011070501; 5 ♂, Erdenesogt Soum, Shargaljuut Gol near Salkhit Uul and forested hillside, N46.47630, E101.43793, 2414 m, 5.vii.2011, S. Podenas, MAIS 2011070504. Bulgan Aimag: 1 ♂, Namnan ul Mountains, 23 km NW Somon Chutag, 1150 m, 21.vii.1968, Exp. Dr. Z. Kaszab, Nr. 1137 ( HNHM); 77 ♂, 6 ♀, Bulgan Soum, Urd Tamir Gol ~ 38 km SW Tsetserleg, 12–13.vii.2004, N47.28244, E101.18793, 1872 m, SRP 04071201; 3 ♂, 1 ♀, Bulgan Soum, Urd Tamir Gol ~ 50 km SW Tsetserleg, N47.21109, E101.07947, 1962 m, 13.vii.2004, SRPT, SRP 04071301; 22 ♂, Bulgan Soum, Urd Tamir Gol braid upstream of bridge ~ 63 km SW Tsetserleg, N47.11192, E101.01048, 2066 m, 13–15.vii.2004, SRPT, SRP 04071302; 1 ♂, 1 ♀, Bulgan Soum, unnamed hillside trib. of Khairkhan Davaani Gol, N46.94286, E100.85532, 2311 m, 13.vii.2004, J.K. Gelhaus, J.C. Morse, A.E.Z. Short, SRP 04071303; 1 ♂, Bulgan Soum, Chandman Gol, trib. of Khairkhan Davaani Gol ~ 80 km SW of Tsetserleg, N47.00226, E100.91051, 2228 m, 13.vii.2004, SRPT, SRP 04071305; 21 ♂, Baruun Tsuuts Gol, 49.4 km NE Hangal, N49.71502, E104.64275, 745 m, 6–7.vii.2005, SRPT, SRP 05070602; 1 ♂, Manhtai Cold Spring, 5 km SW SRP 05070602, N49.6809, E104.56469, 864 m, 7.vii.2005, SRPT, SRP 05070701; 7 ♂, Khyalgant Soum, Baruun Burkhiin Gol, 36.56 km NNW Hangal, N49.62431, E104.24514, 1015 m, 7–8.vii.2005, SRPT, SRP 05070702; 1 ♂, Teshig Soum, West Branch Tariakhtain Gol, N49.70189, E103.80034, 1432 m, 9.vii.2005, SRPT, SRP 05070901; 3 ♂, Teshig Soum, Egiin Gol, 12.7 km SW Teshig, N49.88872, E102.50978, 998 m, 11–13.vii.2005, SRPT, SRP 05071101; 10 ♂, Khishig-undur Soum, Airhan Nuur, South Shore, N49.63456, E102.64865, 937 m, 13.vii.2005, SRPT, SRP 05071301; 6 ♂, Khutag-Ondor Soum, Unit Gol (local name Tolboriin Gol), 20.7 km ESE Khutag-Ondor, N49.3336, E102.97771, 951 m, 25–26.vii.2005, SRPT, SRP 05072501; 6 ♂, Bugat Soum, Altat Gol, 35.2 km SW Khutag-Ondor, N49.21674, E103.10725, 1020 m, 26.vii.2005, SRPT, SRP 05072601; 3 ♂, 12 km S Orchon, N48.53974, E103.60954, 26.vi.2006, 1129 m, SRP 06062602; 2 ♂, Birch ( Betula )-larch ( Larix ) forest, 15 km NW Bulgan, 1435 m, N48.89933, E103.36507, 26.vi.2006, J.K. Gelhaus #1048; 1 ♂, Bulgan Soum, Khar Nuur, ~ 50 km NNW Bulgan, N47.24133, E90.75316, 2563 m, 7.vii.2009, MAIST, MAIS 2009070703; 39 ♂, 1 ♀, Bulgan Soum, Northeast outfall river of Khar Nuur (Black Lake), ~ 50 km NNW Bulgan, N47.24368, E90.75253, 2560 m, 7.vii.2009. Darkhan Aimag: 7 ♂, Shariingol Soum, Khuitnii Gol, N49.46450, E106.66871, 852 m, 21.vii.2003, SRPT, SRP 03072103. Hovsgol Aimag: 1 ♂, Hovsgol Lake, Borsog Gol (river) at shoreline, ~ 1650 m, 3.vii.1996, E. Hunter #96031; 1 ♂, west shore Khovsgol Nuur (lake), Khar-us cold springs, 1700 m, 50°56′30″N, 100°14′37″E, 4.vii.1996, E. Hunter #96066; 4 ♂, 1 ♀, Khovsgol Nuur (lake) at Khankh (town), north end, 1719 m, 51°30′23.11″N, 100°40′19.52″E, 12.vii.1996, E. Hunter #K9612; 1 ♂, Khovsgol Nuur (lake) at Khankh (town), 1719 m, ~51°33′28.30″N, 100°46′41.92″E, 15.vii.1996, E. Hunter #K9615; 1 ♂, Khovsgol Nuur (lake) at Khatgal (dock), 1657 m, 50°27′45″N, 100°10′20″E, 18.vii.1996, E. Hunter #K9618; 1 ♂, 2 ♀, N Somon Chatgal SW shore of Lake Chövsgöl nuur, 1650 m, Exp. Dr. Z. Kaszab, Nr. 1124, 18.viiI. 1968 ( HNHM); 1 ♀, 6 km WNW Somon Tosoncengel, 1480 m, Exp. Dr. Z. Kaszab, Nr. 980, 18.vi.–20.vii.1968 ( HNHM); 26 ♂, Arbulag Soum, Ovor Gylaadin Gol (pooled stream), 7.0 km SE Sumber, 1987 m, N50.12844, E99.60749, 27–28.vi.2006, J.K. Gelhaus #1051, SRP 06062701; 3 ♂, 1 ♀, Ulaan-Uul Soum, unnamed trib. (stream) of Beltes Gol, 34.0 km NE Bayanzurh, 2113 m, N50.42980, E99.21690, 28.vi.2006, J.K. Gelhaus #1054-A, SRP 06062803; 1 ♂, Ulaan Uul Soum, Bagtag Gol (river) and wetland ponds/marshy meadow, 15 km NNE Ulaan Uul, 1598 m, N50.80484, E99.32228, 29–30.vi.2006, J.K. Gelhaus #1057, SRP 06062904; 1 ♂, Renchinlhumbe Soum, Springs and spring fed brooks at Ikh Turuugiin Gol (stream), 35 km NNE Ulaan Uul, 1578 m, N50.99269, E99.35406, 30.vi.–1.vii.2006, J.K. Gelhaus #1058, SRP 06063002; 1 ♂, Renchinlhumbe Soum, Arsayn Gol (river), 16 km N Renchinlhumbe, 1575 m, N51.25339, E99.66685, 2.vii.2006, J.K. Gelhaus #1062, SRP 06070201; 2 ♂, Renchinlhumbe Soum, Jaray (Jarin) Gol (river), 33 km N Renchinlhumbe, 1580 m, N51.39865, E99.75060, 2–3.vii.2006, J.K. Gelhaus #1064, SRP 06070203; 4 ♂, Tsagaannuur Soum, unnamed trib. of Shishged Gol ~ 16 km NW Tsagaannuur (town), 1606 m, N51.45833, E99.20051, 3–4.vii.2006, J.K. Gelhaus #1066, SRP 06070302; 1 ♂, Renchinlhumbe Soum, Larch ( Larix siberica ) forest, 41 km S Tsagaannuur (town), 1604 m, N50.98796, E99.23112, 4.vii.2006, J.K. Gelhaus #1071; 13 ♂, 1 ♀, Ulaan-Uul Soum, Guna (Gunain), Gol (river), 11 km NNW Ulaan-Uul, N50.77518, E99.18892, 1615 m, 4.vii.2006, Malaise trap, SRP 06070403; 1 ♂, Bayanzurh Soum, Beltes Gol (river), 10 km E Bayanzurh, 1688 m, N50.20129, E99.08625, 5.vii.2006, J.K. Gelhaus #1072; 12 ♂, Bayanzurh Soum, Beltes Gol (river), 6 km ENE Bayanzurh, 1682 m, N50.19880, E99.03148, 6.vii.2006, SRP 06070601; 3 ♂, Bayanzurh Soum, Emtin Gol (stream), ~ 2 km S Bayanzurh, 1625 m, N50.15020, E99.97738, 6.vii.2006, J.K. Gelhaus #1075, SRP 06070602; 1 ♂, Moron Soum, Tunamal Nuur (lake), 5.5 km W Arbulag, 1871 m, N49.89920, E99.39433, 6–7.vii.2006, J.K. Gelhaus #1076, SRP 06070603; 4 ♂, Ikh Uul Soum, Selenge Moron (river) ~ 8 km W Ikh-Uul; 1069 m, N49.44641, E101.33625, 7–8.vii.2006, J.K. Gelhaus #1077, SRP 06070701; 1 ♂, Khankh Soum, Sevsuul River, N51.1623611, E100.753833, 1656 m, 25.vi.2005, SRPT; 7 ♂, Khankh Soum, Shagnuul River, N51°15′18.6″, E100°50′45.8″, 1728 m, 25.vi.2005, SRPT; 5 ♂, Khankh Soum, Turag River, N51°17′11.1″, E100°49′12.3″, 1665 m, 25–26.vi.2005, SRPT; 7 ♂, Khankh Soum, Turag River, N51°17′12.6″, E100°49′49.0″, 1663 m, 26.vi.2005, SRPT; 1 ♂, Khankh Soum, Turag River, N51°17′28.1″, E100°48′49.8″, 1668 m, 26.vi.2005, SRPT; 3 ♂, Khankh Soum, Turag River, N51°19′33.7″, E100°54′05.7″, 1795 m, 26.vi.2005, SRPT; 1 ♂, 1 ♀, Khankh Soum, Shagnuul River and Hovsgol Lake nearby, N51.263, E100.779083, 1657 m, 27.vi.2005, SRPT; 3 ♂, Khankh Soum, Shagnuul Gol, 1694 m, 27.vi.2005, SRPT; 5 ♂, Khankh Soum, Shagnuul Gol (middle site), N51.2570278, E100.852444, 1726 m, 27.vi.2005, SRPT; 1 ♂, Khankh Soum, Sevsuul Gol and shore of Hovsgol Lake, N51°09′42.0″, E100°43′42.2″, 1658 m, 28.vi.2005, SRPT; 1 ♂, Khankh Soum, Noyon Gol (upper site), N51.2233611, E100.860833, 1801 m, 28.vi.2005, SRPT; 3 ♂, Khankh Soum, Sevsuul Gol, N51°09′37.1″, E100°45′31.3″, 1666 m, 29.vi.2005, SRPT; 38 ♂, 6 ♀, Erdene Bulgan Soum, Emt Gol, 33.2 km NW Tarialyn, N49.87742, E101.82073, 1496 m, 13–14.vi.2005, SRPT, SRP 05071302; 7 ♂, 9 ♀, Erdene Bulgan Soum, 12.6 km S Erdene Bulgan, N49.99842, E101.60806, 1344 m, 14.vii.2005, SRPT, SRP 05071401; 3 ♂, Erdene Bulgan Soum, Uur (Uuriin) Gol, 32.6 km NE Erdene Bulgan, N50.30193, E101.92869, 1063 m, 14–15.vii.2005, SRPT, SRP 05071403; 1 ♂, 3 ♀, Erdene Bulgan Soum, Zerleg Gol, 35.6 km NE Erdene Bulgan, N50.32719, E101.95058, 1087 m, 15.vii.2005, SRPT, SRP 05071501; 11 ♂, 1 ♀, Steppe outside Tsagaan-Uur (town), N50.53689; E101.52015, 1153 m, 16.vii.2005, J.K. Gelhaus #1016; 67 ♂, 4 ♀, Tsagaan-Uur Soum, Uilgan Gol, 26 km ENE Tsagaan-Uur, N50.6231, E101.87375, 1152 m, 15–16.vii.2005, SRPT, SRP 05071502; 15 ♂, Tsagaan-Uur Soum, Uur (Uuriin) Gol, 7.1 km N Tsagaan-Uur, N50.60052, E101.52344, 1130 m, 16–17.vii.2005, SRPT, SRP 05071601; 32 ♂, Chandmani-Ondor Soum, Darkhunt Gol, 23.5 km E Chandmani-Ondor, N50.50854, E101.26581, 1192 m, 17.vii.2005, SRPT, SRP 05071701; 100 ♂, 2 ♀, Chandmani-Ondor Soum, Khalkhan Gol, 19.8 km E Chandmani-Ondor, N50.51523, E101.21074, elev. 1220 m, 17–18.vii.2005, SRPT, SRP 05071702; 11 ♂, Chandmani-Ondor Soum, unnamed trib. of Hohoo Gol, N50.66022, E100.73886, 1566 m, 18.vii.2005, SRPT, SRP 05071802; 5 ♂, 1 ♀, Chandmani-Ondor Soum, west branch (unnamed) of Khalkhan Gol, upstream of Bolnain Rashaan, 58 km NE Hatgal, N50.78323, E100.79572, 1676 m, 18–19.vii.2005, SRPT, SRP 05071803; 1 ♂, east trib. of upper Hohoo Gol (Bugat Gol), 44.5 km NE Hatgal, N50.69174, E100.66691, 1595 m, 19.vii.2005, SRPT, SRP 05071902; 2 ♂, 1 ♀, Bayanzurkh Soum, Altargana Gol, 4.2 km N Bayanzurkh, N50.21297, E98.96367, 1662 m, 21–22.vii.2005, SRPT, SRP 05072103; 1 ♂, pool near Selenge Moron Gol at Teel (beginning of Selenge below confluence of five rivers), N49.26764, E100.82688, 1182 m, 24–25.vii.2005, SRPT, SRP 05072401; 6 ♂, cold spring on Southern trib. of Beltesiin Gol, 10.1 km SW Sumber, N50.09674, E99.42659, 1882 m, 22.vii.2005, SRPT, SRP 05072202; 4 ♂, Tumurbulag Soum, N49.23175, E99.90027, 1790 m, 23.vii.2005, SRPT, SRP 05072302. Khentii Aimag: 5 ♂, Batshireet Soum, Eg Gol, jct Onon Gol, 8 km NE Batshireet, N48.71888, E110.27771, 1080 m, 16.vii.2011, S. Podenas, MAIS 2011071602; 2 ♂, Dadal Soum, pine forest above Balj Gol, 4.1 km N Dadal, N49.05075, E111.58395, 956 m, 19.vii.2011, S. Podenas; 16 ♂, 3 ♀, Tsenkhermandal Soum, spring headwaters Tsenkher Gol, 29 km N Tsenkhermandal, N47.92564, E109.16114, 1589 m, 23.vii.2011, S. Podenas, MAIS 2011072303. Ovorhangay Aimag: 2 ♂, Kharkhorin Soum, Sant Uul Pass, 23 km SE Shinebrigade, N46.87219, E103.14081, 2007 m, 5.vii.2004, SRPT, SRP 04070501; 1 ♂, Hujirt Soum, Khavtsaliin Gol valley, N46.76292, E103.02561, 1750 m, 5–6.vii.2004, SRPT, SRP 04070504; 4 ♂, Bat-Olziy Soum\Orkhon Gol ~ 34 km W Khujirt, N46.89303, E102.39457, 1610 m, 6.vii.2004, SRPT, SRP 04070602; 9 ♂, Bat-Olziy Soum, braid of Orkhon Gol ~ 40 km W Khujirt, N46.88586, E102.34386, 1646 m, 6–7.vii.2004, SRPT, SRP 04070603; 27 ♂, Bat-Olziy Soum, Orkhon’s Waterfall on Ulaan Gol ~ 300 m S Orkhon Gol, ~ 84 km W Khujirt, N46.78742, E101.96021, 1809 m, 7.vii.2004, SRPT, SRP 04070701; 58 ♂, Bat-Olziy Soum, Ulaan Gol ~ 93 km W Khujirt, N46.73093, E101.88583, 1894 m, 7–8.vii.2004, SRPT, SRP 04070702; 8 ♂, Bat-Olziy Soum, trib. of Ulaan Gol ~ 92 km W Khujirt, N46.72826, E101.89317, 1941 m, 8.vii.2004, SRPT, SRP 04070801; 1 ♂, Kharkhorin Soum, Kharzani Gol 3.5 km NE Shankh Bridgade, N47.05722, E102.99191, 1519 m, 8.vii.2004, SRPT, SRP 04070803; 8 ♂, Uyanga Soum, Khangai Mountains, Khuysiyn Naiman Nuur Strictly Protected Area, streams entering east side of Shireet Nuur (lake), down from davaa (pass), N46.51908, E101.84940, 2446 m, 7.vii.2011, C. R. Nelson & T.A. McKnight & MAIST, MAIS 2011070701; 9 ♂, Uyanga Soum, Khangai Mountains, Shuranga Gol, just outside entrance to Khuysiyn Naiman Nuur Strictly Protected Area, N46.47528, E101.88175, 2451 m, 7.vii.2011, S. Podenas, MAIS 2011070703; 10 ♂, Uyanga Soum, Ovtiyn Gol, 20 km SE Bat-Olziy, N46.64265, E102.30885, 2069 m, 8.vii.2011, S. Podenas, MAIS 2011070801; 9 ♂, Bat-Olziy Soum, Ovtiyn Gol, 10 km SE Bat-Olziy, N46.75972, E102.30905, 1847 m, 8.vii.2011, S. Podenas, MAIS 2011070803. Selenge Aimag: 12 ♂, 2 ♀, Mandal Soum, Boyant Gol (trib. of Tunkhel Gol), 13.8 km above town Tunkhel, 2.0 km up- stream of Shonkhor Camp, N48.67237, E106.87851, 1078 m, 16–17.vii.2003, SRPT, SRP 03071602; 2 ♂, Bugant/ Yaroo Soum, Bar Chuluu Gol, N49.03282, E106.96935, 975 m, 17.vii.2003, SRPT, SRP 03071704; 5 ♂, Bugant/Yaroo Soum, Khongiin Gol at Yeroo Gol confluence, N49.08636, E107.30750, 943 m, 18–19.vii.2003, SRPT, SRP 03071803; 32 ♂, Bugant/Yaroo Soum, unnamed trib. of Yalbag Gol, 49.7 km W Zuunkharaa Bridge, N49°04′57.7″, E106°53′51.7″, 1115 m, 17.vii.2003, Suvdtsetseg, SRP 03071703; 6 ♂, Khuder Soum, Galttiin Gol c. 6 km S Khuder, N49.73769, E107.48766, 702 m, 24–25.vii.2003, SRPT, SRP 03072402; 8 ♂, Khuder Soum, Tsagaan Shiluustiin Gol ca. 5 km S Khuder, N49.73763, E107.49333, 704 m, 23–24.vii.2003, SRPT, SRP 03072302; 5 ♂, Khuder Soum, unnamed stream 54.2 km E Tavin (Yeroo), N49.77751, E107.26517, 907 m, 25.vii.2003, SRPT, SRP 03072501; 2 ♂, Mandal Soum, Dart Gol 38.9 km W Zuunkharaa Bridge, N49.05571, E106.80383, 1228 m, 17.vii.2003, J.K. Gelhaus, S. Podenas, SRP 03071702; 4 ♂, Mandal Soum, Sharingol headwaters 4.6 km below southern ridgetop, N49.10789, E106.65275, 1124 m, 21.vii.2003, SRPT, SRP 03072101; 3 ♂, Pine forest slope, N50°19′53.3″, E105°04′48.9″, 765 m, 26.vii.2003, J.K. Gelhaus #936; 10 ♂, Tsagaannuur Soum, SW shore of Tsagaan Nuur, N49.95588, E105.33791, 668 m, 25.vii.2003, SRPT, SRP 03072502; 1 ♂, Baruunharaa Soum, Haraa Gol, 2.3 km S Bayangol, N48.87738, E106.12375, 807 m, 4–5.vii.2005, SRPT, SRP 05070402; 2 ♂, Zuunburen Soum, Selenge Gol, 7.8 km NW Zuunburen, N50.10285, E105.78217, 647 m, 5–6.vii.2005, SRPT, SRP 05070501; 1 ♂, Tsagaannuur Soum,Wetland, 10.1 km SW Tsagaanuur, N50.04519, E105.36084, 706 m, 6.vii.2005, SRPT, SRP 05070601; 3 ♂, davaa (pass) SW Sogooch, N49.83555, E104.99729, 971 m, 6.vii.2005, C. R. Nelson #8184 & SRPT; 4 ♂, Tsagaannuur Soum, davaa (mountain pass), 53.6 km S Tsagaannuur (town), N49.83561, E104.99727, 973 m, 6.vii.2005, J.K. Gelhaus #995. Tov Aimag: 2 ♂, Erdene Soum, Gorkhi Terelj NP, Terelj Gol braid down-stream from Terelj, 10.6 km N Mungut Rock Road, N47.97944, E107.47761, 1487 m, 5.vii.2003, SRPT, SRP 03070502; 7 ♂, Erdene Soum, Gorkhi Terelj NP; Tuul River ca. 100 m upstream of confluence with Terelj River, N47.96806, E107.59365, 1467 m, 5.vii.2003, SRPT, SRP 03070503; 4 ♂, Erdene Soum, Gorkhi Terelj NP, W Bayangin Gol 8.2 km upstream of Tuul River Road, N48.08026, E107.63841, 1527 m, 6.vii.2003, SRPT, SRP 03070603; 3 ♂, Erdene Soum, Gorkhi Terelj NP, Khokh Chuluutiin Gol, N48.12675, E107.57996, 1680 m, 6.vii.2003, SRPT, SRP 03070604; 2 ♂, Erdene Soum, Gorkhi Terelj NP, W Bayangin Gol, 20.1 km upstream of Tuul River Road, N48.16668, E107.69548, 1623 m, 6.vii.2003, SRPT, SRP 03070605; 2 ♂, Tov Aimag Erdene Soum, Gorkhi Terelj NP, E Bayangin Gol, 12.9 km upstream of Tuul River Bridge, N48.14846, E107.75838, 1596 m, 7.vii.2003, SRPT, SRP 03070703; 1 ♂, Erdene Soum, Gorkhi Terelj NP, E Bayangin Gol, 4.2 km upstream of Tuul River Bridge, N48.07956, E107.77729, 1526 m, 7.vii.2003, SRPT, SRP 03070704; 1 ♂, Erdene Soum, Gorkhi Terelj NP, Tuul River, 10.1 km upstream of Tuul River Bridge, N48.09549, E107.84265, 1531 m, 8.vii.2003, SRPT, SRP 03070801; 17 ♂, Erdene Soum, Gorkhi Terelj NP, unnamed braid of Tuul River, N48.14846, E107.91267, 1564 m, 8–9.vii.2003, SRPT, SRP 03070805; 9 ♂, 2 ♀, Erdene Soum, Gorkhi Terelj NP, unnamed trib. of Tuul River on W side, 1.6 km upstream from Daichin crossing, N48.21780, E107.90392, 1594 m, 9.vii.2003, SRPT, SRP 03070902; 1 ♂, 1 ♀, Erdene Soum, Gorkhi Terelj NP, unnamed trib. of Tuul River on W side, c. 5 km upstream from Daichin crossing, N48.24734, E107.90589, 1610 m, 10.vii.2003, SRPT, SRP 03071001; 5 ♂, 1 ♀, Erdene Soum, Gorkhi Terelj NP, Tuul River at Daichin crossing, N48°12′41.6″, E107°54′78.0″, 10.vii.2003, Namkhaidorj, Monkhbayar, SRP 03071003; 8 ♂, Erdene Soum, Gorkhi Terelj NP, unnamed trib. of Tuul River on E side 8.5 km downstream of Galtain Gol, N48.09720, E107.84928, 1542 m, 11.vii.2003, SRPT, SRP 03071101; 4 ♂, Erdene Soum, Gorkhi Terelj NP, E side oxbow of Tuul River, N48°06′53.6″, E107°53′35.5″, 1560 m, 11.vii.2003, SRPT, SRP 03071102; 14 ♂, Batsumber Soum, unnamed trib. of Segnogor Gol, 14.3 km upstream of Segnogor Amralt, N48.43755, E107.03066, 1255 m, 15.vii.2003, SRPT, SRP 03071501; 11 ♂, Erdene Soum, unnamed river NE Khagiin Har Nuur, Gorkhi Terelj NP, N48.42080, E107.91406, 1818 m, 28.vii.2003, SRPT, SRP 03072802; 36 ♂, 1 ♀, Cherlen Rver basin, Dund burchijn gol, 1487 m, 2–3.viii.2003, N48.4438889, E108.624, S. Podenas; 8 ♂, Cherlen River basin, Terelzhijn gol, 1482 m, 4–5.viii.2003, N48.4227778, E108.6505, S. Podenas; 8 ♂, Cherlen gol, 1393 m, 5–6.viii.2003, N48.3744444, E108.857222, S. Podenas; 2 ♂, 1 ♀, Zuud baidlagyn gol, 1437 m, 6–7.viii.2003, N48.1925, E108.6116, S. Podenas; 4 ♂, Zuunmod Soum, Turgeniy Gol (stream), 8 km NNW Zuunmod, 1581 m, N47.77758, E106.92780, 14.vii.2006, J.K. Gelhaus #1073-B, SRP 06071401; 5 ♂, Odlog Udleg canyon, Batsumber Soum, N48.26480, E106.93345, 1359 m, 16.vii.2006, SRP 06071602; 1 ♂, Batsumber Soum, Bayan gol, N48.35418, E106.8383, 1145 m, 5.vii.2011, O. Yadamsuren, MAIS 2011070502; 1 ♂, Erdene Soum, Baruun Bayan gol, N48.07718, E107.63927, 1544 m, 6.vii.2011, O. Yadamsuren, MAIS 2011070603; 2 ♂, Mongomorit Soum, Tuul gol, N48.04708, E107.74532, 1517 m, 7.vii.2011, O. Yadamsuren, MAIS 2011070701; 5 ♂, Erdene Soum, Galtain gol, N48.13485, E107.92422, 1565 m, 7.vii.2011, O. Yadamsuren, MAIS 2011070703; 1 ♂, Erdene Soum, Tuul gol, N48.18108, E107.91869, 1573 m, 7.vii.2011, O. Yadamsuren, MAIS 2011070704; 2 ♂, Erdene Soum, Gorkhi Terelj NP, stream 0.5 km N Gorkhi Davaa (Pass), N47.94981, E107.45511, 1593 m, 11.vii.2011, S. Podenas, MAIS 2011071101; 30 ♂, 2 ♀, Erdene Soum, Bulnayn Gol, jct Galttyn Gol, 33 km W Mongonmorit, N48.12563, E108.02163, 1596 m, 12.vii.2011, S. Podenas, MAIS 2011071204; 5 ♂, Erdene Soum, Shar Bulag, Yellow Spring, 26 km W Mongonmorit, N48.14814, E108.07956, 1629 m, 13.vii.2011, S. Podenas, MAIS 2011071301; 13 ♂, Erdene Soum, Zuun Baydlag Gol, 15 km SW Mongonmorit, N48.17390, E108.26500, 1641 m, 14.vii.2011, S. Podenas, MAIS 2011071401; 12 ♂, 2 ♀, Mongonmorit Soum, Zuun Baydlag Gol, downstream, 15 km SW Mongonmorit, N48.10818, E108.36736, 1475 m, 14.vii.2011, S. Podenas, MAIS 2011071402; 8 ♂, Mongonmorit Soum, Burkhin Gol, 4.5km NW Mongonmorit, N48.25508, E108.48447, 1475m, 14.vii.2011,S.Podenas, MAIS 2011071403; 8 ♂, 1 ♀, Mongonmorit Soum, Kherlen Gol, 16 km SE Mongonmorit, N48.13212, E108.63913, 1376 m, 15.vii.2011, S. Podenas, MAIS 2011071501; 1 ♂, Mongonmorit Soum, Zuun Baydlag Gol, 4 km SW Mongonmorit, N48.09511, E108.38592, 1452 m, 25.vii.2011, MAIS 2011072501. Uvs Aimag: 1 ♀, 4 km ESE Pass Ulaan davaa, between Lake Örög nuur and Ulaangom, 1700 m, Exp. Dr. Z. Kaszab, Nr. 1074, 6.vii.1968 ( HNHM); 1 ♂, Tarialan Soum, Kharkhiraa Gol, 4 km W Tarialan, N49.77959, E91.86026, 1453 m, 2–3.vii.2010, MAIST, MAIS 2010070201; 1 ♂, Khovd Soum, Goojuur Gol, 25 km NE Khovd, N49.42978, E91.19503, 1975 m, 5–6.vii.2010, MAIST, MAIS 2010070505; 3 ♂, Khovd Soum, Goojuur Waterfall, 29 km NE Khovd, N49.44144, E91.25202, 2274 m, 6.vii.2010, MAIST, MAIS 2010070601; 1 ♂, Khovd Soum, braid of Shiver Gol, 12 km E Khovd, N49.24592, E91.06149, 1489 m, 6– 7.vii.2010, MAIST, MAIS 2010070604; 1 ♂, Bokhmoron Soum, Altan Gadasni Gol, 7 km N Jigertei Bag, Gulzat Protected Area, N49.99679, E90.27887, 1763 m, 10–11.vii.2010, MAIST, MAIS 2010071003; 1 ♂, Turgen Soum, Khondlon Gol, 4.5 km W Turgen, N50.08403, E91.61472, 1295 m, 13–14.vii.2010, MAIST, MAIS 2010071303; 2 ♂, Turgen Soum, Springs S side of Khondlon Gol, 6 km W Turgen, N50.07458, E91.60140, 1316 m, 14.vii.2010, MAIST, MAIS 2010071401; 13 ♂, 5 ♀, Davst Soum, Tokhilog Gol, by Mongolian military border outpost, N50.69165, E92.59283, 1003 m, 14–15.vii.2010, MAIST, MAIS 2010071403; 16 ♂, Sagil Soum, Borshoo Gol, 29 km NNE Sagil, N50.57615, E91.77029, 1281 m, 15–16.vii.2010, MAIST, MAIS 2010071502; 2 ♂, Turgen Soum, Javartain Gol, 33.5 km SW Turgen, N49.88653, E91.34708, 1871 m, 16.vii.2010, MAIST, MAIS 2010071603; 2 ♂, Turgen Soum, Javartain Gol & Turgen Gol, 33 km SW Turgen, N49.89234, E91.35239, 1849 m, 16–17.vii.2010, MAIST, MAIS 2010071602; 12 ♂, Ondorkhangai Soum, Baruunturuuni Gol & Morguugiin Gol, 20 km NNW Ondorkhangai, N49.44592, E94.79595, 1688 m, 19–20.vii.2010, MAIST, MAIS 2010071902; 10 ♂, 1 ♀, Tsagaankhayrkhan Soum, Onchun Gol, 15 km ESE Tsagaankhayrkhan, N49.38851, E94.44290, 1832 m, 20– 21.vii.2010, MAIST, MAIS 2010072001. Zavkhan Aimag: 1 ♂, Choit chunch, 24 km E Somon Songino, 2000 m, Exp. Dr. Z. Kaszab, Nr. 1096, 12.vii.1986 ( HNHM); 1 ♂, 24 km E Somon Songino, 2000 m, Exp. Dr. Z. Kaszab, Nr. 1097, 12.vii.1968 ( HNHM); 1 ♂, Ikh-Uul Soum, Ideriin Gol ~ 28 km E Tosontsengel, N48.71968, E98.65184, 1654 m, 19–20.vii.2004, SRPT, SRP 04071903; 1 ♂, Tosontsengel Soum, Delgarakhiin Gol ~ 16 km S Tosontsengel, N48.61518, E98.23096, 1761 m, 20–21.vii.2004, SRPT, SRP 04072001; 20 ♂, 1 ♀, Tosontsengel Soum, Gunza Gol ~ 38 km SW Tosontsengel, N48.62624, E97.89617, 1831 m, 21–22.vii.2004, SRPT, SRP 04072102; 6 ♂, 1 ♀, Mountain pass (davaa), N48.42180, E97.83063, 2135 m, 22.vii.2004, J.K. Gelhaus #979; 9 ♂, 2 ♀, Tosontsengel Soum, Tegshiin Gol ~ 70 km SW Tosontsengel, N48.34410, E97.87312, 1927 m, 22.vii.2004, SRPT, SRP 04072201; 2 ♂, Tosontsengel Soum, unnamed reed lake and marsh by Tegshiin Gol, ~ 70 km SW Tosontsengel, N48°19′52.6″, E97°51′52.2″, 22.vii.2004, A.E.Z. Short, J.K. Gelhaus, Enkhnasan, SRP 04072202; 7 ♂, Ider Soum Ideriin Gol ~ 4 km NE Zuunmod/Ider, N48.24995, E97.40627, 1929 m, 22–23.vii.2004, SRPT, SRP 04072203; 15 ♂, 1 ♀, Ider Soum, Dogshin/Nogoon Nuur ~ 21 km SE Zuunmod/Ider, N48.06257, E97.55064, 2054 m, 23–24.vii.2004, SRPT, SRP 04072302; 1 ♂, Ider Soum, Ideriin Gol ~ 22 km SE Zuunmod/Ider, N48.05989, E97.55073, 2053 m, 23– 24.vii.2004, SRPT, SRP 04072303; 1 ♂, Ider Soum, Zagastain Gol ~ 11 km NNE Zagastain Davaa, N48.15779, E97.21441, 2108 m, 24.vii.2004, SRPT, SRP 04072401; 6 ♂, Telmen Soum, Ideriin Gol ~ 15 km SSW Telmen/ Ovogdii, N48.53255, E97.52093, 1823 m, 24–25.vii.2004, SRPT, SRP 04072402; 1 ♂, Tosontsengel Soum, Ar Teeliin Gol ~ 24 km SW Tosontsengel, N48.65028, E98.36208, 1803 m, 25.vii.2004, SRPT, SRP 04072501; 48 ♂, 5 ♀, Ikh-Uul Soum, Deed Tsetsuukhiin Gol ~ 10 km NW Solongotiin Davaa, N48.36436, E98.92737, 2072 m, 25.vii.2004, SRPT, SRP 04072502.

Elevation range in Mongolia. Adults were collected at altitudes ranging from 640 to 2800 m.

Period of activity. Adults are active from mid-June through to early August.

Known distribution. Kazakhstan, Mongolia ( Map 14 ) and Russia (western and eastern parts of Siberia, Far East).

Redescription. Male. Body length 12.0– 16.1 mm, wing length 13.9–18.0 mm. General body coloration brownish yellow.

Head. Gray. Vertex and occiput gray with indistinct median line. Rostrum brownish yellow, dorsally sparsely dusted with gray, nasus conspicuous. Antenna 13-segmented, if bent backward reaching beyond base of wing. Scape, pedicel and first flagellomere yellow, succeeding flagellomere brown. Each flagellomere except first one with weak basal enlargement and deeply incised. Apical flagellomere very small, reduced, distinctly shorter than preceding flagellomere. Verticils longer than corresponding segments. Palpus dark brown.

Thorax. Brownish gray. Pronotum light brown with broad brown median line. Prescutum and presutural scutum gray with 4 brownish longitudinal stripes indistinctly bordered by brown. Intermediate pair separated. Postsutural scutum gray, scutal lobe with 2 spots indistinctly bordered by brown. Scutellum light brown with brown median line. Postnotum gray pruinose with brown median line. Pleura pruinose, brownish gray. Coxa yellowish, dusted with gray. Trochanters yellowish. Femora yellowish with narrowly darkened distal part. Tibiae yellowish with darkened distal part. Tarsal segments brown. Tarsal claws without tooth. Wing patterned with brown. Halter with pale stem and brown knob.

Abdomen. Yellow, with conspicuous broad dark brown dorsal and lateral stripes. Segment 1 sparsely dusted with gray. Lateral and caudal borders of tergites pale.

Hypopygium. Yellow. Tergite 9 forming narrowed concave plate ( Fig. 163 ). Main body of tergal plate pale with blackened rim. Posterior margin of tergal plate broadly emarginated. Lateral angles of tergal plate obtuse and blackened. Anterior portion of tergal plate raised into broad sclerotised border. Border medially interrupted by pale. Gonocoxite shaped as large subrectangular plate, apex gently bifid with large obtuse black tooth and small denticle ( Fig. 164 ). Outer gonostylus widestest at middle with apex rounded ( Fig. 165 ). Inner gonostylus large, mid-dorsal edge with blackened plate, beak sclerotised and rounded ( Fig. 166 ). Gonocoxal fragment ( Fig. 167 ) with medial sclerites slender, flattened posteriorly, anteriorly fused by membrane and together forming V-shaped structure. Lateral sclerite very small and inconspicuous. Aedeagal guide in shape of triangular groove, gonapophyses short and wrinkled, adminicular rods long ( Fig. 168 ). Sperm pump with central vesicle flattened ( Fig. 169 ). Compressor apodeme shaped as large plate without median incision. Posterior immovable apodeme much longer than compressor apodeme appearing as slender lobe. about same length as compressor apodeme lobe with rounded apex.Anterior immovable apodeme laterally obtuse. Aedeagus shaped as elongate tube, about 6.9 times as long as sperm pump, basally and medially dark brown, apically passing into yellow. Distal part with lateral and ventral incisions shorter than in T. (V.) virgatula , appearing trident-shaped ( Fig. 170 ).

Female. Body length 20.9–26.0 mm, wing length 2.0–3.0 mm ( Fig. 3 ). Female differs from male by reduced wings and elongate abdomen.Antenna short, if bent backward reaching base of wing. Scape, pedicel and first flagellomere yellowish, succeeding flagellomeres brownish with small weak basal enlargements.

Ovipositor ( Figs 171–173 ). Tergite 10 shiny brown. Cercus brown, longer than tergite 10, relatively broad with tip obtuse, outer margin with indistinct small serration ( Fig. 171 ). Sternite 8 anteriorly with 2 small lateral projections. Hypovalva extremely short, sclerotized and stout projection, distinctly shorter than in T. (V.) virgatula ( Fig. 172 ). Sternite 9 with anterior part distinctly broadened, posterior part short and acute. Furca anteriorly dark brown and narrow, posteriorly membranous except median yellowish-brown area ( Fig. 173 ). Bursa copulatrix and spermathecal ducts without sclerotisation, generally similar to T. (V.) virgatula . Spermatheca spherical ( Fig. 14 ).

Figures

FIGURES 1–3. Adults of Mongolian T. (Vestiplex), general view. 1. T. (Vestiplex) subcentralis, male habitus, left lateral view; 2. T. (Vestiplex) mediovittata, female habitus, left lateral view; 3. T. (Vestiplex) longitudinalis, female habitus, left lateral view.

FIGURES 4–9. Gonocoxal fragments of Mongolian T. (Vestiplex), epandrium and hypandrium removed, dorsal view. 4. T. (Vestiplex) longitudinalis; 5. T. (Vestiplex) mediovittata; 6. T. (Vestiplex) scripta; 7. T. (Vestiplex) pallitergata; 8. T. (V). balioptera; 9. T. (Vestiplex) leucoprocta. Abbreviations: aa, anterior apodeme of medial sclerite of gonocoxal fragment; pa, posterior apodeme of medial sclerite of gonocoxal fragment; b, base of medial sclerite of gonocoxal fragment; gcx, gonocoxite; ig, inner gonostylus; il, inner lobe of lateral sclerite of gonocoxal fragment; ls, lateral sclerite of gonocoxal fragment; ms, medial sclerite of gonocoxal fragment; ol, outer lobe of lateral sclerite of gonocoxal fragment. Scales of 4–6, 8–9 = 0.5 mm, 7 = 0.25 mm.

FIGURES 14–27. Spermathecae of Mongolian T. (Vestiplex). 14. T. (Vestiplex) longitudinalis; 15. T. (Vestiplex) virgatula; 16. T. (Vestiplex) pallitergata; 17. T. (Vestiplex) laccata; 18. T. (Vestiplex) balioptera; 19. T. (Vestiplex) nubeculosa; 20. T. (Vestiplex) scripta; 21. T. (Vestiplex) kiritshenkoi; 22. T. (Vestiplex) mediovittata; 23. T. (Vestiplex) subcentralis; 24. T. (Vestiplex) sintenisi; 25. T. (Vestiplex) coronifera; 26. T. (Vestiplex) tchukchi; 27. T. (Vestiplex) leucoprocta.

FIGURES 163–170. Hypopygium of male T. (Vestiplex) longitudinalis. 163. Tergite 9, dorsal view; 164. Gonocoxite, lateral view; 165. Left outer gonostylus, lateral view; 166. Left inner gonostylus, lateral view; 167. Gonocoxal fragment, dorsal view; 168. Aedeagal guide, lateral view; 169. Sperm pump, dorsal view; 170. Distal part of aedeagus, lateral view. Scale of 163–169 = 0.5 mm, 170 = 0.25 mm.

FIGURES 171–173. Ovipositor of female T. (Vestiplex) longitudinalis. 171. Ovipositor, left lateral view; 172. Sternite 8 with hypovalvae, ventral view; 173. Sternite 9, furca and bursa copulatrix, dorsal view. Scale of 171 = 0.7 mm, 172–173 = 0.5 mm.

MAPS 13–14. Distribution maps of T. (Vestiplex) crane flies in Mongolia. 13. T. (Vestiplex) longitudinalis; 14. T. (Vestiplex) virgatula.