Nanotrachia, INTERMEDIA (SOLEM, 1984)

Köhler, Frank & Criscione, Francesco, 2013, Small snails in a big place: a radiation in the semi-arid rangelands in northern Australia (Eupulmonata, Camaenidae, Nanotrachia gen. nov.), Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 169 (1), pp. 103-123 : 119

NANOTRACHIA INTERMEDIA ( SOLEM, 1984)

Ordtrachia intermedia Solem, 1984: 666–669 , pl. 56, figures 172–173.

Material examined: Table 1.

Diagnosis

Shell ( Fig. 4A ): Average in size; periphery shouldered, weakly keeled; spire moderately elevated; umbilicus open, wide, sometimes partly concealed by columellar reflection; protoconch and teleoconch with dense, pointed periostracal projections and fine axial growth lines; teleoconch with blunt, regular, widely spaced axial ribs; apertural lip reflected, moderately expanded; horn.

Genitalia ( Fig. 6A, B ): Penial retractor muscle inserting on vas–epiphallic junction. Epiphallus about as long as penis, thick, tapering in thickness towards Taxonomic remarks

Type species of Nanotrachia gen. nov., not congeneric with other species originally assigned to the genus Ordtrachia , as revealed by the molecular phylogeny ( Fig. 2 ).

Figures

Figure 2. Bayesian phylogram based on analyses of concatenated COI and 16S sequences. Nodal support (%): Bayesian posterior clade probabilities (BPP, above branches), ML bootstrap (BTSP, below branches); only BPP values ± 90% and BTSP values ± 70% are shown; asterisks = 100%.

Figure 4. Shells of Nanotrachia species studied herein. A, N. intermedia (Solem, 1984) (paratype AM C.146110); B, Nanotrachia carinata sp. nov. (paratype WAM S49100); C, N. coronata sp. nov. (paratype WAM S49086); D, N. costulata costulata sp. nov. (WAM S49088); E, N. levis sp. nov. (paratype WAM S49096); F, N. orientalis (Solem, 1984) (paratype AM C.146105); G, Mouldingia occidentalis (Solem, 1984) (paratype AM C.146107). Scale bar = 1 cm.

Figure 6. Penial anatomy. A, Nanotrachia intermedia (Solem, 1984) (WAM S49195); B, N. intermedia (Solem, 1984) (WAM S49194); C, Nanotrachia carinata sp. nov. (holotype WAM S66300); D, N. levis sp. nov. (holotype WAM S6630). Insets showing schematic cross section of mid-penis. Scale bar = 2 mm.